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EN
Cratopetalura petruleviciusi gen. et sp. nov. is the third genus and species of the Mesozoic petalurid family Cretapetaluridae from the Lower Cretaceous of Brazil. With the recent discovery of another representative of this family in the Lower Cretaceous of England, it demonstrates the great diversity of this group during this period.
EN
Mantodea are rather scarce in the fossil record, especially those belonging to the mantise crown group. Four fossil mantids are described from the lowermost Eocene amber of Oise (France), two Chaeteessidae considered as “genus and species incertae sedis”, and two Mantoididae, described as a new genus and species Pseudomantoida extendidera. We also describe a new specimen of Arvernineura insignis from the Paleocene of Menat (France), confirming the attribution of this taxon to the Chaeteessidae. These fossils are of great interest for future dating of the crown group Mantodea, being the oldest Chaeteessidae and Mantoididae. We propose a new genus name Louispitonia nom. nov. in replacement of Archaeophlebia Piton, 1940 preoccupied by Archaeophlebia Ris, 1909 (Odonata) with Archaeophlebia enigmatica as its type species.
EN
A new family Vosegidae is described for Vosegus triassicus gen. et sp. nov. from the Anisian (early Middle Triassic) Grès à Voltzia Formation of the Vosges (north−eastern France). This is the oldest Aphidomorpha presenting the combination of characters as: elongate and tapered pterostigma with straight posterior margin, thick common stem of veins Sc+R+(M)+CuA, arcuate RP separated well basad of pterostigma base, three−branched M with free base, CuA forked, veins not differentiated in their thickness. The morphological characters presented by Vosegus are spread among the different extinct and extant lineages of Aphidomorpha, but the combination presented is unique. The relationships of this new family to other Triassic Sternorrhyncha are discussed, concluding its placement in Aphidomorpha: Triassoaphidoidea. The specificity of Grès à Voltzia fossil site in respect to early evolution of aphids is presented, with two competing explanations for size diminution, considering the Early Triassic biotic recovery versus the correlation between early aphid diversification and the exploitation of new niches, new host plants and habitats. This new finding increases the taxonomic diversity of the Grès à Voltzia fauna, expanding our knowledge of Early Triassic Aphidomorpha diversity and evolution.
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Content available A new Chinese Mesozoic stick insect
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EN
Renphasma sinica gen. et sp. nov. is described from the Early Cretaceous Yixian Formation in Liaoning. It has the general habitus of the Mesozoic “stick−insects”, with a short thorax, and broad and long tegmina, as long as the hindwings. Its pattern of tegmina venation is typical of the Mesozoic Phasmatodea, also present in the Eocene stick insect family Gallophasmatidae, viz. archaeorthopteran organisation of median and cubital veins. On the other side it has a vomer typical of modern Phasmatodea, both supporting the hypothesis that the Mesozoic, Cainozoic, and modern stick insects belong to the same clade. The pattern of dark and hyaline spots on the wings of Renphasma is probably related to mimicry with plants.
EN
The new fossil thrips Uzelothrips eocenicus P. Nel and A. Nel sp. nov. (Thysanoptera: Uzelothripidae) is described from two lowermost Eocene amber−preserved specimens (one macropterous and one apterous). The family Uzelothripidae is only known so far from a single extant species, Uzelothrips scabrosus. The fossils differ from the extant species only by the antennal segments III and IV, which appear distinctly separated instead of being fused as in the in the extant U. scabrosus. Dark−coloured hyphae and conidia of the Dothideomycetes (Ascomycota) which are likely to belong to the sooty moulds (Capnodiales) are attached to the apterous fossil specimen. We consider this arthropod−fungus association not to be accidental since these fungi are also found in extant specimens of these uzelothripids, suggesting very specific long−term interactions and strong habitat specificity.
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EN
The enigmatic Svalbard Palaeocene fossil taxon Hymenopterites deperditus is revised, and is neither a wasp nor a plant seed, but turns out to be a bug hemelytra corresponding to the oldest described reduviid bug. It can be attributable to the “emesine−saicine clade”. The presence in the Svalbard Palaeocene of this insectivorous bug, showing possible mimicry of a wasp model, confirms the presence of diverse entomofauna and of rather warm palaeoclimatic conditions.
EN
Two braconid wasp taxa with enlarged eyes and ocelli indicative of probable nocturnal activity are discussed and described from the lowermost Eocene amber of the Paris Basin.The new tribe Palaeocharmontini nov., for new genus Palaeocharmon with type species Palaeocharmon basalis sp. nov. is described and illustrated in the subfamily Charmontinae. The similarity of this taxon with members of subfamilies Helconinae, Homolobinae, and Brachistinae is shown and the peculiar character [presence of hind wing recurrent vein (m−cu)] unknown previously in non−cyclostome braconids is discussed. A third fossil species of the genus Phanerotoma (Ph. menieri sp. nov.) is described and compared with known species from the Baltic amber.
EN
A new interpretation of the wing venation of the fossil insect order Caloneurodea is proposed. Complex fusions of veins are demonstrated, supporting the assignment of the Caloneurodea to the Panorthoptera, i.e. the order is closely related to the Orthoptera. A new diagnosis of the order is proposed. The following taxa are re-described after new interpretation, material or preparation, or are newly described: Caloneura dawsoni Brongniart, 1885 (= Boltonaloneura subtilis (Bolton, 1925) syn. nov.), Gigagramma carpenteri gen. and sp. nov.; Sthenarocera pachytyloides Brongniart, 1885, Pruvostiella lecomtei (Pruvost, 1919), Apsidoneura flexa Carpenter, 1943, Homaloptila similis (Meunier, 1911), Pleisiogramma medialis Carpenter, 1943 (=Pleisiogramma reducta Carpenter, 1943 syn. nov.), Paleuthygramma acuta Carpenter, 1943, Paleuthygramma sharovi sp. nov., Euthygramma parallelum Martynov, 1928 (= Euthygramma curvatum Martynov, 1931 syn. nov.; = Paleuthygrammella aberrans (Martynov, 1938) syn. nov.), Anomalogramma parva Carpenter, 1943, Nanogramma gandi gen. and sp. nov., and Ligogramma sinuosa gen. and sp. nov. The family Gelasopteridae Carpenter, 1976 is revised and its ordinal relationships are discussed. After the proposed phylogenetic frame, mutation(s) occurring in homeotic genes are expected to be responsible of the unusual wing morphology of the Caloneurodea, mixing fore- and hind-wing 'panorthopteriď features.
EN
The new genus and species Alboconis cretacica (oldest known Aleuropteryginae: Fontenelleini) and the coniopterygine new genus and species Gallosemidalis eocenica, are described, respectively from a late Albian and an early Eocene French amber. From Lebanese amber, the early Cretaceous Aleuropteryginae Libanoconis fadiacra (Whalley, 1980) is refigured and discussed.
EN
Pseudoxyelocerus bascharagensis gen. et sp. nov., the oldest representative of the Tenthredinoidea and Xyelotomidae, based of a single forewing, and an enigmatic “Symphyta” family incertae sedis, based on a hindwing, are described from the Toarcian of Luxembourg. The relationships of the genera currently included in Xyelotomidae are briefly discussed. The genera Undatoma, Liaotoma, Leridatoma, and Davidsmithia have the unique apomorphy in the wing venation of the Tenthredinoidea minus Xyelotomidae. The Xyelotomidae is probably a paraphyletic family. Only a phylogenetic analysis will help to solve these problems. There is no evidence supporting the previous assignments of the fossil genera Vitimilarva and Kuengilarva to the family Xyelotomidae.
EN
The first fossil genus and species of Tetratomidae (Coleoptera) is described, from the Lower Cretaceous amber deposits of France. The new genus represents the first insect to be recognized inside an opaque piece of amber, through the use of propagation phase contrast X-ray microtomography using synchrotron radiation. This new finding proves the capabilities of this imaging technique in amber inclusions, as well as increases the knowledge of fossil tenebrionoids, a group scarcely recognized in the Cretaceous fossil record.
EN
The position of the Palaeozoic Anthracoptilidae has been a major problem of insect systematics for over a century. The previous hypotheses suggested affinities of Anthracoptilidae with the Palaeodictyopteroida, Protorthoptera, Hypoperlida, stem-Mantodea, Paraneoptera, Holometabola, or Eoblattida. Herein we put forward a new hypothesis based mainly on a comprehensive re-evaluation of the wing venation characters and re-examination of the type material of Anthracoptilus perrieri and Mesoptilus dolloi. The Anthracoptilidae are considered as belonging to the paoliid lineage, sister group of the Dictyoptera. In particular this result refutes the presence of Mantodea in the Paleozoic fossil record. The two families Strephocladidae and Strephoneuridae, are considered as junior synonyms of the Anthracoptilidae, while the previous synonymy of the Anthracoptilidae with the Ischnoneuridae is rejected. We consider the Permarrhaphidae, previously considered as synonym of the Anthracoptilidae, (and Permarrhaphus) as Insecta incertae sedis. The following new taxa are proposed: Mesoptilus carpenteri sp. nov. from the early Permian of Wellington Formation in Oklahoma (USA) which extends the range and distribution of the genus; Pseudomesoptilus gen. nov. is designated to include Mesoptilus sellardsi; Strephocladus permianus sp. nov. from the middle Permian of South of France; Westphaloptilus gallicus gen. et sp. nov. from the Bashkirian of the North of France.
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