The behavior and ultrastructure of the endosperm haustorium was studied in the natural tetraploid Trifolium pratense L. (Leguminosae). The lower portion of the endosperm generally remains free-nuclear and functions as an aggressive haustorium. The haustorium contains dense cytoplasm with many free nuclei, which usually aggregate in groups of 2-4. The haustorium presumably transports metabolites from the adjacent tissues to the developing endosperm proper. It is functional up to the late heart-shaped stage of embryo development.
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