The Ociesęki Sandstone Formation represents the Holmia-Schmidtiellus and the Protolenus-Issafeniella zones. Seven ichnospecies of Rusophycus, including Rusophycus exsilius isp. nov, are recognized in this formation. Moreover, Rusophycus ispp. A, B, C are recognized. The diagnoses for R. dispar, R. crebrus and R. magnus have been emended and the stratigraphic range of R. versans has been extended. The presence of Rusophycus dispar may suggest a palaeozoogeographical connection between the Małopolska Block, the Baltica palaeocontinent and the Laurentia palaeocontinent.
In general, the trace fossil Rusophycus, preserved as a concave-upward structure on the top of a bed, is considered to be a fossilized marking, made by a trace maker. The structures described from the Cambrian (Furongian) of central Poland are genetically related to Rusophycus. However, despite their occurrence on the tops of beds, they are not fossilized traces, but compaction-related features, resulting from differential sandstone and mudstone compaction with possible mediation by organic-rich, heterolithic sediments. The preservation of these structures probably was influenced by biofilms or biomats.
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