Eoiocossus validas gen. and sp. nov., a new genus and species belonging to Palaeontinidae (Insecta, Hemiptera), is described from the Middle Jurassic of Daohugou, Inner Mongolia, China. It differs from other genera as follows: large forewing with small clavus, Sc terminating in costal margin beyond the nodus, nodal line along m₄ -cua partly, CuA₂ with two branches and A₂ developed. The ripple-like posterior margin and fresh colour pattern of Eoiocossus may be sex characteristics. The marginal membrane and ambient vein have not distinct evolutionary implications for the Palaeontinidae.
Some structural characters and morphometric variables – size, body shape and proportions, wing shape and structure – that appear in insects to be linked with flight performance, are discussed and evaluated, and methods are described for deriving these from fossil material. Some wing design categories associated with particular flight techniques and capabilities are identified. Their use in reconstructing the flight performance of extinct insects is illustrated with reference to Carboniferous palaeodictyopteroids and Mesozoic palaeontinoid Hemiptera.
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