Menathyris wilsoni gen. et sp. nov. is described from the Limestone and Marl Member (Early Ladinian) of the Saharonim Formation (early Anisian–late Carnian), at Har Gevanim, Makhtesh Ramon, southern Israel. The Saharonim Formation represents the main transgressive phase of the Middle Triassic. The subunit, from which the shells were collected, contains the molluscs Neoschizodus laevigatus (Ziethen), Myophoriopsis cf. subundata (Schauroth), Lima cf. tellei Bittner, Pecten discites (Schlotheim), Ostrea montiscaprilis Klipstein, Pseudoplacunopsis fissistriata (Winkler), Gevanites awadi (Parnes), and the conodont Pseudofurnishius murcianus (Van den Boogaard), the last-mentioned indicative of the Fassanian (Early Ladinian).
Erythrosuchidae is a clade of early archosauriform reptiles that were large-bodied, hypercarnivorous, possibly apex predators in late Early and Middle Triassic ecosystems following the Permo-Triassic mass extinction. Chalishevia cothurnata from the late Middle Triassic (Ladinian) of Russia, is the stratigraphically youngest known erythrosuchid species, but the holotype and referred material of this taxon has received little study. Here, we provide the first detailed anatomical description of C. cothurnata, including comparisons to other erythrosuchids. Although known from relatively fragmentary material, the anatomy of C. cothurnata is distinctive, including an autapomorphic strongly slanted ventral border of the antorbital fossa. The presence of a large accessory opening (the “accessory antorbital fenestra”) in the skull between the premaxilla, nasal and maxilla, together with the inferred presence of a narrow postnarial process of the premaxilla that articulated with a slot on the nasal, provides strong evidence for a sister taxon relationship between C. cothurnata and the erythrosuchid Shansisuchus shansisuchus from the early Middle Triassic (Anisian) of China. The inferred basal skull length of C. cothurnata was approximately 80 cm, making it one of the largest erythrosuchids known.
Prestosuchus chiniquensis is the most famous “rauisuchian” described by Friedrich von Huene, eight decades ago, and several specimens have been assigned to this taxon since then. In the present contribution, we provide the first detailed description of a complete and very well preserved skull (including the braincase) assigned to Prestosuchus chiniquensis from the Dinodontosaurus Assemblage Zone of the Santa Maria Supersequence of southern Brazil. The detailed description of the skull of Prestosuchus chiniquensis, besides increasing the knowledge about this taxon, may help elucidate the taxonomic relationships of pseudosuchians even further, since most of the characters used in phylogenetic analyzes are cranial. The presence of the subnarial fenestra, a controvertial extra opening on the skull of “rauisuchians”, is thoroughly discussed considering the evidence provided by this new specimen. We consider that the small slit-opening between the premaxilla and the maxilla in Prestosuchus chiniquensis, can not safely be considered a true fenestra, but indicates more likely the existence of some degree of cranial kinesis between these elements which can result in different relative positions of the bones after definitive burial and fossilization, so that the size and shape of this opening is taphonomically controlled. Complementarily, the digital reconstruction of its cranial endocast was developed both from the observation of the preserved braincase and from CT scan images, which resulted in obtaining the first endocast known for a “rauisuchian”. The endocast of Prestosuchus chiniquensis exhibited some remarkable convergences with that of theropod dinosaurs, which could be a reflection of the similar niches they occupied, since “rauisuchians” were the top predators at the end of the Late Triassic, before the extinction of all non-crocodylomorph pseudosuchians.
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The late Ladinian to Late Triassic succession of the Tahtalidag (upper) Nappe of the Antalya nappes was studied in the Egregindere section, north of the city of Antalya, SW Turkey. The chert bands in the central part of the section have yielded poorly to moderately preserved radiolarians documenting the Late Ladinian Muelleritortis firma and Muelleritortis cochleata radiolarian zones. Based on the Egregindere succession, a major deepening event, evidenced by radiolarian cherts, took place between the middle and late Late Ladinian. The Late Triassic thick-bedded neritic limestones represent a shallowing-upward sequence, which formed as a result of the horst-like rising of the Tahtalidag Nappe during the Late Triassic block faulting. Fifty-nine radiolarian taxa have been determined from the Upper Ladinian of the Egregindere section. One species (Muelleritortis elegans) and two subspecies (Muelleritortis firma equispinosa and Muelleritortis firma globosa) are described as new.