Great thicknesses of sand and greywacke were deposited on the margins of megacontinents during the Ediacaran Period (620–542 Ma). Zircon age populations in sediments with long deep-time flat profiles distinguish passive margin sedimentation from shorter humped zircon profiles characteristic of sediments derived from volcanic arcs and their feeder zones in active margins. An example of a single hump detrital profile is given by an Ediacaran Period volcano present in the Charnian Supergroup in the Anglo-Brabant Massif of the East Avalonia terrane. This Gondwana fragment was originally part of the West Africa craton and was subsequently accreted to Laurentia. A volcanic complex with sediments carrying an Ediacaran biota is overlain by Triassic sediments. The main phase of eruption at c. 561 Ma provides a single hump zircon age histogram with a few pre-eruption zircon xenocrysts up to 40 Ma older
Global palaeogeographic maps were constructed for eight time intervals in the Palaeozoic. The maps contain information concerning plate tectonics and palaeoenvironment during the Cambrian, Ordovician, Silurian, Devonian and Carboniferous. The East European Craton belonged to the Palaeozoic Baltica Plate, which originated as a result of disintegration of the supercontinent Pannotia during the early Cambrian. Baltica included part of Poland and adjacent areas northeast of a line that extends between Scania and the Black Sea. This plate was located in the Southern Hemisphere and drifted northward during Early Palaeozoic time. The Early Ordovician was the time of maximum dispersion of continents during the Palaeozoic. Avalonia probably started to drift away from Gondwana and moved towards Baltica during Ordovician time. Between Gondwana, Baltica, Avalonia and Laurentia, a large longitudinal oceanic unit, known as the Rheic Ocean, was formed. Avalonia was probably sutured to Baltica by the end of the Ordovician or in the Early Silurian. This process was dominated by the strike-slip suturing of the two continents, rather than a full-scale continent-continent collision. Silurian was a time of Caledonian orogeny, closing of the Early Palaeozoic oceans, collision of Baltica with Avalonia and Laurentia and the assembly of the supercontinent Laurussia. The Variscan orogeny in Poland was caused by the collision of the Bohemian Massif plates and the Protocarpathian terrane with Laurussia. The Protocarpathian terrane acted as an indentor that caused thrust tectonics in the East European Platform, Holy Cross Mountains and the Lublin area.
The faunal history of insects in the various fragments of Gondwana is presented. The first part of the paper summarizes the current knowledge of its insect-bearing localities, particularly their stratigraphy and fossil content, emphasizing the record of the higher systematic groups. The second part discusses some trends of their palaeobiodiversity as evidenced from the above mentioned sites. Generally, the knowledge of the fossil Gondwanan insect faunae is still much lower than that of the Laurasian ones, but has considerably increased over the last decade. Altogether about 85 localities are known from Gondwana, with a maximum of sites in Permian and a minimum in Jurassic times. Best represented is South America. Fossil insects of Gondwana are probably less known than those of Laurasia due to inadequate exploration rather than unfavourable conditions for the formation of deposits.
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Analiza stabilnych izotopów tlenu i węgla w wapieniach środkowego i górnego ordowiku regionu kieleckiego Gór Świętokrzyskich wskazuje na zmiany ich składu w czasie. Największe wahania (w skali PDB) występują w wyższym ordowiku (górny karadok) i pojawiają się po okresie aktywności prądów wstępujących. Fluktuacje te byty przypuszczalnie spowodowane czynnikami klimatycznymi, kontrolowanymi przez zlodowacenie na Gondwanie, które spowodowały zachwianie stratyfikacji gęstościowej i termicznej oceanów (Wilde & Berry, 1984; Railsback i in., 1990; Wilde, 1991). Wartości [delta]^18O i [delta]^13C są zbliżone do wyników uzyskanych dla kopalnych, chłodno- lub umiarkowanie ciepłowodnych osadów węglanowych oraz mieszczą się w przedziale wartości stwierdzonych przez wielu badaczy dla ordowiku.
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Oxygen and carbon isotopic studies were carried on bulk-rock samples from the Middle and Upper Ordovician limestones taken j'rom the Mójcza section in Kielce Region of the Holy Cross Mts. Measurements of [delta]^18O and [delta]^13C are close to results obtained from ancient temperate- and warm-temperate water carbonates. The greatest changes both [delta]^18O and [delta]^13C are associated with upper part of the Mójcza section (Upper Caradoc) and appear after ceasing of upwelling currents activity. Carbon and oxygen isotopic fluctuations are concurrent with changes in conodont assemblages (Dzik, 1999) and were probably caused by climatic factor, controlled by glaciation on Gondwana.
A diverse microvertebrate fauna is described from the Virgin Hills and Napier formations, Bugle Gap Limestone Canning Basin, Western Australia. Measured sections at Horse Spring and Casey Falls (Virgin Hills Formation) and South Oscar Range (Napier Formation) comprise proximal to distal slope carbonates ranging in age from the Late Devonian Frasnian to middle Famennian. A total of 18 chondrichthyan taxa are identified based on teeth, including the first record of Thrinacodus tranquillus, Cladoides wildungensis, Protacrodus serra and Lissodus lusavorichi from the Canning Basin. A new species, Diademodus dominicus sp. nov. is also described and provides the first record of this genus outside of Laurussia. In addition, the upper range of Australolepis seddoni has been extended to Late Devonian conodont Zone 11, making it the youngest known occurrence for this species. The Virgin Hills and Napier formations microvertebrate faunas show close affinities to faunas recovered from other areas of Gondwana, including eastern Australia, Iran, Morocco and South China, which is consistent with known conodont and trilobite faunas of the same age.
The study of the strophomenide brachiopods of the subfamily Rafinesquininae present in the main Upper Ordovician sections, representing the Mediterranean margin of Gondwana, has revealed an increase in diversity of the group at the region during that time. The studied collections are from the Moroccan Anti-Atlas, the Iberian and the Armorican massifs, the Iberian Chains, Pyrenees, Montagne Noire, Sardinia, and Bohemia. Two genera of the subfamily Rafinesquininae have been recorded. Of them, the cosmopolitan Rafinesquina is the only one previously reported from the region and Kjaerina is found for the first time outside Avalonia, Baltica, and Laurentia. Additionally, two new subgenera have been described, Kjaerina (Villasina) and Rafinesquina (Mesogeina). Furthermore, the new species Rafinesquina (Mesogeina) gabianensis, Rafinesquina (Mesogeina) loredensis, Kjaerina (Kjaerina) gondwanensis, Kjaerina (Villasina) pedronaensis, Kjaerina (Villasina) pyrenaica, and Kjaerina (Villasina) meloui have been described. In addition, other species of these genera previously known from isolated localities in the region, such as Rafinesquina pseudoloricata, Rafinesquina pomoides, and Hedstroemina almadenensis are revised and their geographic range expanded. The adaptive radiation experienced by the rafinesquinines at the Mediterranean region during middle to late Katian, was probably related to changes in the regime of sedimentation and water temperature caused by the global warming Boda event.
We report here the discovery of a Cretaceous mammal from the “Red Sandstone Group” of southwestern Tanzania. This specimen is one of only a very few Cretaceous mammals known from Gondwana in general and Africa in particular. The specimen consists of a short, deep left dentary that bore a large, procumbent central incisor, and five single−rooted, hypsodont cheek−teeth. The specimen is very tentatively identified as a sudamericid, and thus may represent the first African record of an enigmatic clade of mammals, the Gondwanatheria, which is otherwise known from the Late Cretaceous and Paleogene of several other Gondwanan landmasses. Unfortunately, the precise age of the specimen could not be determined. If it is pre−Campanian and if its identity as a sudamercid is corroborated through subsequent discoveries, it represents the earliest known gondwanatherian. If the specimen is from the Campanian or Maastrichtian, and again assuming its identification is correct, it has the potential to refute a recently formulated biogeographic hypothesis predicting the absence of certain terrestrial and freshwater vertebrate taxa, including gondwanatherians, in Africa (i.e., those that evolved elsewhere on Gondwana after Africa became an isolated landmass).
A review of the Cambrian edrioasteroid echinoderm Cambraster cannati is made based on new collections from the Iberian Chains (NE Spain) and Montagne Noire (France). New morphological data include a completely articulated oral area and details of ambulacra. Specimens ranging from 4 to 26 mm in diameter provide detailed information concerning the full ontogeny. Important changes through ontogeny mainly affect the marginal ring and the plating pattern of the aboral surface. Comparison with other species of Cambrasterindicates that the aboral surface of Cambraster tastudorum from Australia shows strong resemblance to juvenile specimens of C. cannati. Cambraster cannati was attached directly to the substrate and inhabited relatively high energy, offshore environments from the west margin of Gondwana. Abnormalities in the skeleton are described for the first time in a Cambrian edrioasteroid.
Sparse fish microremains have been found in marine limestones from the Middle Devonian (Givetian) Skały Formation (Sitka Coral-Crinoid Limestone Member and Sierżawy Member), Świętomarz–Śniadka section, Bodzentyn Syncline, Łysogóry Region, northern Holy Cross Mountains, associated with conodonts of the hemiansatus to ansatus zones. Thelodont scales referred here to Australolepis sp. cf. A. seddoni come from near Śniadka village, from samples dated as hemiansatus to rhenanus/varcus zones. This increases the known range for the genus from its original find in Western Australia. The presence of a thelodont in the late Middle Devonian in Poland extends the known distribution of turiniids around the peri-Gondwana shorelines of Palaeotethys.
In the Balkans, the Serbo-Macedonian Unit (SMU), Serbia, is thrust bounded by the composite Tethyan Vardar Zone and the Carpatho-Balkanides. The SMU actually emerges from beneath the Neoalpine Miocene-Pliocene deposits. Both provenance and geodynamic position of the SMU are poorly known and still debated. This paper reviews the data hitherto published and includes some new field data interpretations. The SMU is composed of a Neoproterozoic-Cambrian high-grade (para- and ortho-) gneiss with peraluminous magmatic arc components (560-470 Ma). The SMU is in the contact with Neoproterozoic upper Ordovician-Carboniferous low-grade metasedimentary succession of an accretionary wedge assembly represented by the Supragetic basement. The SMU basement became folded, sheared and metamorphosed around 490-450 Ma. Paleomagnetic data point to high southern latitudes and a peri-Gondwanan position of the SMU at that time, which concurs with glaciomarine evidence recorded from the upper Ordovician sediments at the base of an accretionary wedge succession. Based on the published data and field survey in the Stalać region, we correlate the SMU with the pre-Mesozoic gneiss terrane exposed in the Strona-Ceneri zone of the Alps. This terrane, identified as the Cenerian orogen of the Alaskan subduction type, developed at an active margin of Gondwana during middle Ordovician times. The SMU basement, with augen and migmatitic gneisses and arc-related peraluminous magmatic bodies, developed at this margin as part of the Cenerian belt or its equivalent. Such an orogenic edifice proved transient and in the earliest Silurian the SMU fragments drifted away being bound for Baltica (amalgamated Moesian microplate and Danubian terrane) to which they became accreted in the Carboniferous and included in the southern European branch of the Variscan orogen (Marginal Dacides/Carpatho-Balkanides). Despite consider-able Variscan and Alpine reworking, the pre-Variscan, Cenerian-type crustal assembly along with an inferred boundary between the magmatic arc and the accretionary wedge, accompanied by back-arc/forearc deposits, are still decipherable in the Western Balkan countries.
Abelisaurid theropods were one of the most diverse groups of predatory dinosaurs in Gondwana during the Cretaceous. The group is characterized by a tall, wide skull and robust cervical region. This morphology is thought to have facilitated specialized feeding behaviors such as prolonged contact with prey. The Late Cretaceous abelisaurid Majungasaurus crenatissimus typifies this abelisaurid cranial morphotype. Recent fossil discoveries of this species include a partial growth series that allows for the first time an investigation of ontogenetic variation in cranial morphology in a representative abelisaurid. Herein we examine growth trajectories in the shape of individual cranial bones and articulated skulls of Majungasaurus using geometric morphometrics. Several major changes in skull shape were observed through ontogeny, including an increase in the height of the jugal, postorbital, and quadratojugal, an increase in the extent of the contacts between bones, and a decrease in the circumference of the orbit. The skull transitions from relatively short in the smallest individual to tall and robust in large adults, as is seen in other theropods. Such morphological change during ontogeny would likely have resulted in different biomechanical properties and feeding behaviors between small and large individuals. These findings provide a post-hatching developmental framework for understanding the evolution of the distinctive tall skull morphology seen in abelisaurids and other large-sized theropod dinosaurs.
A microfauna of small shelly fossils (SSF) is reported here for the first time from middle Cambrian (Series 3, Stage 5) subsurface strata of the Torgau-Doberlug Syncline (TDS), Central Germany. Considering that this microfauna is strongly limited and poorly preserved the material is quite abundant and diverse. The assemblage consists of molluscs (pelagiellids, bivalves), coeloscleritophorans (chancelloriids, halkieriids), poriferids, protoconodonts, cambroclaves, hyoliths, brachiopods, and disarticulated echinoderm remains. Additionally, a probable pterobranch hemichordate is noted. The assemblage is dominated by epifaunal suspension feeders from mid- to outer shelf depositional settings. Stratigraphically it represents (together with rare trilobites) the oldest middle Cambrian (Series 3, Stage 5) fauna known from Central Germany and the entire Saxothuringian Zone. Regardless the taphonomic problems related to the SSF occurrence, close palaeobiogeographic relations are indicated with the Mediterranean shelf of West Gondwana (especially with the areas of southwestern Europe and Morocco). The reported microfauna coupled with recent trilobite and palynomorph research supports assumptions that the Cambrian succession in the TDS is by far more complete than hitherto suggested, emphasizing its importance as a region yielding Cambrian rocks in Central Europe.