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EN
During the last two decades, an abundant selachian assemblage has been collected from the late Ypresian (NP12) fossiliferous sands of Prémontré (Aisne, northern France) but has received little attention. Sharks of the family Triakidae (Carcharhiniformes) are particularly well represented and all are described and figured herein. Among them, two new species of the genus Galeorhinus are described: G. duchaussoisi sp. nov. and G. louisi sp. nov.; these are compared to the common Paleogene G. ypresiensis which is refigured. Another triakid taxon, the genus Gomphogaleus gen. nov., is described. Most of the triakids have been recorded elsewhere in the North Atlantic region, suggesting a wider distribution than expected for these small sharks during the Paleogene. The present paper updates the list of selachians from Prémontré, bringing the number of taxa from 19 to 33 (including 22 sharks and 11 batoids) and improving our knowledge of the ancient North Atlantic Ypresian selachian fauna. Despite this vastly improved record, it is clear that fossil data are still very incomplete and insufficient for calibrating phylogenetic hypotheses of living forms. Review of the Prémontré fauna shows that the Triakidae were much more diverse and broadly distributed than at present, suggesting that the limited distribution and low diversity of living forms is probably a recent phenomenon.
EN
Fossil deep-sea selachians are rare and their diversity underestimated as a consequence of the scarcity of available outcrops of sediments containing them. Here we report a new fossil locality from the Middle Eocene of southwestern France and give a first synthesis of the deep-sea deposits of this area which have yielded one of the richest fossil selachian faunas ever to have inhabited the continental slope. The fossil records of deep-sea sharks and rays are discussed in the context of these new fossil occurrences, a literature review and the recent phylogenetic hypothesis.
EN
New or so far poorly known neoselachians from the Cenomanian and Turonian of SW France are described. The material studied herein comes from nine localities in the Charentes region, comprising palaeoenvironments ranging from coastal to open marine environments, and consists of two orectolobiforms, six lamniforms, and four rajiforms. The new taxa are Squalicorax coquandi sp. nov. and Roulletia bureaui gen. et sp. nov. within lamniforms, and Hamrabatis bernardezi sp. nov., Archingeayia sistaci gen. et sp. nov., and Engolismaia couillardi gen. et sp. nov. within rajiforms. New specimens of Odontaspis rochebrunei Sauvage, 1880 from the type area allow redescription of this taxon, assigned herein to the genus Cenocarcharias. Occurrences of Squalicorax baharijensis, S. cf. intermedius, and Archaeolamna sp., previously unrecorded from this region, and Almascyllium, a genus generally described from younger strata, are also noted, and improve knowledge of mid−Cretaceous selachian faunas from Western Europe.
EN
We describe a new primitive proboscidean, Daouitherium rebouli gen. et sp. nov., from the early Ypresian of the Ouled Abdoun Basin, Morocco, which also yielded Phosphatherium. It is the earliest known large mammal from Africa and one of the oldest known proboscideans. It has true lophodont molars similar to those of Barytherium and Numidotherium. It is closer to these genera and more advanced than Phosphatherium (e.g., morphology of the mandible), but it is also primitive in striking features known also in Phosphatherium (absence of diastema, retention of two additional teeth in front of p2). A parsimony analysis of Daouitherium suggests its intermediate phylogenetic position between the basal, small Phosphatherium and the large, more derived Numidotherium and Barytherium. Daouitherium is a better candidate for the ancestry of N. koholense than Phosphatherium, but it is also specialized. Daouitherium and Numidotherium may belong to the same basal radiation of “Barytherioidea”. However, the family referral of Daouitherium is uncertain (Numidotheriidae?). The discovery of such a large and derived proboscidean with respect to Phosphatherium in the same African beds of such antiquity is evidence of an unexpected early diversity of proboscideans and of the old origin of the order. It also supports the African origin of Proboscidea s.s.
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