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EN
Upper Cretaceous to lower Palaeogene carbonate and siliciclastic deposits that crop out widely in the Haymana and Polatlı districts (Ankara Province) of the Haymana Basin (Central Anatolia) are rich in larger benthic foraminifera, various macrofossils and ichnofossils. The ichnofossils of the Haymana and Yeşilyurt formations were studied at five localities. The Upper Cretaceous siliciclastics of the Haymana Formation contain moderately diverse trace fossils, belonging to the deep-sea Nereites ichnofacies. The Paleocene siliciclastic and carbonate deposits of the Yeşilyurt Formation comprise similar trace fossils, which do not show significant changes in comparison to the Haymana Formation. This indicates that the K-Pg boundary extinction event did not affect the ichnofauna with any longer consequences. The Upper Cretaceous coarser, siliciclastic deposits of the Haymana and Beyobası formations are rich in shallow-marine, larger benthic foraminifera, including species of Orbitoides, Omphalocyclus, Siderolites, Hellenocyclina, and Loftusia, whereas fine siliciclastic deposits contain abundant planktonic, open-marine foraminifera, such as Globotruncana and Heterohelix. Palaeogene siliciclastic to carbonate deposits of the Kartal, Yeşilyurt and Çaldağ formations are rich in the larger, benthic foraminifera Nummulites, Discocyclina, Assilina and Alveolina. The larger foraminifers have been redeposited from nearby, shallow-marine parts of the basin.
EN
Palynological investigation of the Vrabchov dol locality (Western Bulgaria) which recently yielded fragmentary dinosaur bones attributed to the clade Titanosauria, reveals well-preserved sporomorph assemblages dominated by angiosperm pollen from the Normapolles group, spores and rare gymnosperms. The age assessment of the studied sequence is based on the diagnostic Normapolles species, such as Oculopollis orbicularis Góczán, 1964, Oculopollis zaklinskaiae Góczán, 1964, Krutzschipollis spatiosus Góczán in Góczán et al., 1967 and Krutzschipollis crassus (Góczán, 1964) Góczán in Góczán et al., 1967. The concurrent presence of these pollen species suggests a late Santonian–early Campanian age for the succession. The sporomorph association is encountered in a palynofacies dominated by continental elements, including translucent phytoclasts (tissues, wood remains and plant cuticles). The sedimentary succession shows no evidence of marine elements and a very low proportion of AOM that attests to deposition within a lagoonal to foreshore marine environment, with high continental input and short transportation. The vegetation in the studied area was primarily composed of a range of Normapolles-producing angiosperms and secondarily of pteridophyte spore-producing plants. Gymnosperms were rare. Such a vegetation pattern reflects a warm, seasonally dry climate during the late Santonian–earliest Campanian in the studied area. The dinosaurs inhabited a wet lowland area, probably rich in herbaceous plants.
EN
A new cephalopod collection from the Campanian-Maastrichtian boundary interval of NE Mexico, consisting of 1076 individuals assigned to 29 species and 22 genera is presented. This collection is a mix of ammonoids, one coleoid and one nautilid, which originate from at least three ammonoid biozones: The upper Campanian Exiteloceras jenneyi and Nostoceras (Nostoceras) hyatti zones, and the lower Maastrichtian Pachydiscus (Pachydiscus) neubergicus Zone. The age of the collection is thus middle late Campanian to late early Maastrichtian, and it closes a stratigraphic gap between faunas described formerly from this region. The specimens are nuclei collected from the desert pavement. The abundance of specimens allows for a comparison to other Campanian-Maastrichtian ammonoid records from Mexico, North America and Europe.
EN
This taxonomy and stratigraphy of the Upper Campanian and Lower Maastrichtian Belemnitella lineages, from the Middle Vistula sections, based on new collections, is presented. The correlation to the basal Maastrichtian standard GSSP at Tercis, France, is provided based on inoceramid bivalve stratigraphy. The Artificial Neural Networks (ANN), particularly the self-organizing Kohonen algorithm, was applied to taxonomic discrimination. Eight morphotypes within the genus Belemnitella, understood here as natural species populations, were recognised. Five of these are assigned to known taxa: Belemnitella mucronata, B. posterior, B. minor [= B. minor I and B. minor II], B. langei and B. najdini; and three, B. sp. a, B. sp. 1 and B. sp. 2, are left in open nomenclature. Four Belemnitella zones are proposed. Due to its palaeogeographic position, between Western and Eastern Europe, the Middle Vistula section is characterized by the co-occurrence of Belemnitella species from those two areas. Consequently, it enables better correlation of Belemnitella-based schemes; the East European B. najdini and B. posterior are placed next to West European B. minor chronospecies I and II. The Campanian/Maastrichtian boundary, as currently defined, is placed at the top of the najdini – posterior Zone, which is an equivalent of the Belemnella-based boundary, i.e. at the base of the Belemnella obtusa / Belemnella vistulensis zones. Within the top of the najdini – posterior Zone occurs a level (an interval of only a few metres), where nearly all of the Upper Campanian Belemnitella disappear. This level coincides with taxonomic changes observed within the co-occurring representatives of genus Belemnella.
EN
The stratigraphical distribution of Porosphaera globularis, a common calcareous sponge in the Upper Cretaceous (mostly Campanian and Maastrichtian) of Poland was studied. The presented material, both new and from museum collections, comes from the Campanian of the Miechów Synclinorium, in southern Poland, and from the Lower Campanian of Mielnik in the south-eastern part of the Mazury-Podlasie Homocline, in eastern Poland. The significance of the species in extra-regional correlation, its palaeobiogeography and stratigraphical potential is critically reviewed.
EN
The taxonomy and stratigraphy of the Upper Campanian and Lower Maastrichtian belemnites from the Vistula (central Poland) and Kronsmoor (northern Germany) sections are revised on the basis of new collections from the Vistula section as well as a reinvestigation of the classic collection of Schulz from the Kronsmoor section. For the taxonomic description a new biometric procedure is proposed, which can be applied to both the genera Belemnella and Belemnitella. For the species-level taxa recognition the Artificial Neural Networks method, the self-organizing Kohonen algorithm, was implemented. This new taxonomic and methodological approach enabled the recognition of nine species of the genus Belemnella. Five of them can be assigned to the existing species B. lanceolata, B. longissima, B. inflata, B. obtusa and B. vistulensis. However, the species concept differs from that applied by Schulz (1979). As a consequence, the stratigraphic ranges of these species are modified. Four species are left in open nomenclature and represent possibly new species. Future studies may reveal that they might be assigned to East European forms from Ukraine or Russia. The species of Belemnella recognized are placed into the stratigraphic framework based on the standard ammonite and inoceramid bivalve zonations, especially those recognized in the Vistula section. The newly proposed belemnite zonation for the Vistula and Kronsmoor sections is correlated via inoceramids with the standard GSSP at Tercis, France, in order to identify the base of the Maastrichtian Stage. The Campanian/Maastrichtian boundary as defined in Tercis is placed here at the base of the newly defined B. obtusa and B. vistulensis Zones ["obtusa/vistulensis"] - thus it is markedly higher than the traditional boundary based on the FAD of representatives of the genus Belemnella - This new boundary coincides well with a distinct turnover of belemnite guard morphology and represents one of the most important points in the early evolutionary history of Belemnella. Three belemnite zones defined by their lower boundaries are recognized in the Campanian/Maastrichtian interval, in addition to three subzones recognized within the B. obtusa Superzone. The B. lanceolata and B. inflata zones as understood here are referred to the Upper Campanian [Tercis definition]. The B. obtusa Zone is subdivided into three subzones, viz.: Belemenlla vistulensis, Belemnella sp. G and Belemnella sp. F, which are referred to the Lower Maastrichtian [Tercis definition]. The fast evolving species of Belemnella enable the proposal of a biostratigraphic scheme with a resolution that is higher than those based on inoceramid bivalves and ammonites - the longevity of a belemnite zone could be as low as 200Ky.
EN
The formal definition of the Global Stratotype Section and Point (GSSP) for the base of the Maastrichtian Stage at Tercis, southwest France, is based on the first or last occurrences of twelve taxa, including three species of ammonites, Pachydiscus (Pachydiscus) neubergicus, Diplomoceras cylindraceum, and Nostoceras (Nostoceras) hyatti. The taxonomy and stratigraphical distribution of these, and allied forms, are studied, on the basis of material from the opoka succession across the upper Campanian.lowermost Maastrichtian boundary in the Middle Vistula River section, central Poland. in view of the imprecise location of two ammonite GSSP markers in that section, a direct ammonite-based correlation with the GSSP at Tercis is impossible. however, data available indicate that in the Middle Vistula River section the first occurrence of Pachydiscus (P.) neubergicus and the last occurrence of Nostoceras (N.) hyatti are situated significantly higher than the first occurrence of Belemnella lanceolata, the traditional belemnite marker for the base of the Maastrichtian Stage in the Boreal Realm, and that Diplomoceras cylindraceum appears significantly below this level. There are no unequivocal records of Pachydiscus (P.) neubergicus from the Belemnella lanceolata Zone s.l. in the Middle Vistula River section.
EN
Results of detailed multistratigraphic analyses of the Campanian.Maastrichtian boundary section at Kronsmoor in northern Germany are summarised and calibrated with the GSSP at Tercis les Bains, southwest France. Additional markers for the definition of the boundary in the Boreal Realm are proposed, and a detailed carbon isotope curve around the Campanian.Maastrichtian boundary in the chalk facies of the Boreal epicontinental sea is presented. The C isotopic GSSP marker for global correlation is the markedly abrupt decrease of c. 0.7 [per mil] [delta^13]C directly at the Campanian.Maastrichtian boundary as dated by ammonites. In electronic borehole measurements the Kronsmoor section covers the SP peaks 53 to 64 and the base of the Maastrichtian being situated just below SP peak 60. The first occurrence (FO) of the ammonite Pachydiscus neubergicus, which corresponds to biohorizon 1 at Tercis, falls in the upper part of nannofossil Zone UC15, at both localities. Biohorizon 3 is the FO of the ammonite Diplomoceras cylindraceum, which first appears in the Upper Campanian of Tercis and at Kronsmoor enters significantly above the FO of Belemnella lanceolata, the conventional Boreal belemnite marker for the base of the Maastrichtian Stage. Based on ammonite evidence, the internationally accepted base of the Maastrichtian at Kronsmoor is located between the FOs of Diplomoceras cylindraceum (Upper Campanian) and Pachydiscus neubergicus (Lower Maastrichtian) c. 11 m above flint layer F 600, at which the first representatives of the belemnite genus Belemnella, in particular Bn. lanceolata occur. The latter thus is a Late Campanian species, appearing c. 450 ky prior to the ammonite-based boundary. The FOs of Belemnella pseudobtusa (sensu Schulz) resp. Belemnella obtusa (sensu Remin) directly at the boundary can be use as the coleoid proxy for the definition of the base of the Maastrichtian in the Boreal Realm. To define the boundary by benthic foraminifera the last occurrence (LO) of Neoflabellina praereticulata is suitable. Biohorizon 12, as defined at Tercis, involves the nannofossil Uniplanarius trifidus, however, at Kronsmoor this species is rare, occurs only sporadically and also significantly lower in comparison to Tercis. It is possible though to compare and correlate nannofossil events between Kronsmoor and Tercis using cosmopolitan taxa such as Broinsonia parca constricta and Eiffellithus eximius. The LO of the latter appears to be situated just above the boundary in both sections; it follows from this that the top of nannofossil Zone UC15 is of Early Maastrichtian age.
EN
The topmost part of the Oxfordian limestones, building the Zakrzówek Horst in Kraków, is featured by a network of minute fissures, filled with Upper Cretaceous limestones. Fissures are dominantly subhorizontal, anastomosing and polygonal in plane. They are filled with white limestones representing mostly foraminiferal- calcisphere wackestones, with subordinate amount of quartz pebbles and fragments of stromatolite coming from the latest Turonian-?Early Coniacian conglomerate overlying Oxfordian basement. The fissures are seismically- induced injection dykes. In contrast to gravitationally-filled neptunian dykes the recognised injection dykes were filled by overpressured soft sediments. Foraminifera within some dykes are abundant, and dominated by planktonic forms, which indicate the Early/Late Campanian age (Globotruncana ventricosa and Globotruncanita calcarata zones) of the filling, and hence date also the synsedimentary tectonics. Abundant and diversified keeled globo- truncanids in the Campanian of the Kraków region are recognised for the first time. Other important findings at the studied section include karstic cavities featuring the surface of the Oxfordian bedrock filled with conglomerates of the latest Turonian-?Early Coniacian age based on foraminifera and nannoplankton, and lack of Santonian deposits, which elsewhere are common in the Upper Cretaceous sequences in the Kraków region. The discovered Campanian dykes provide new evidence for the Late Cretaceous tectonic activity on the Kraków Swell related to the Subhercynian tectonism, which resulted among others in stratigraphic hiatuses and unconformities characte- ristic of the Turonian-Santonian interval of this area.
PL
W pelagicznych osadach jednostki podśląskiej (marglach węglowieckich, marglach typu frydeckiego, marglach żegocińskich, piaskowcach z Rybia) oraz w marglach bakulitowych jednostki skolskiej i marglach z Bonarki występujących na monoklinie śląsko-krakowskiej (odsłonięcie Bonarka w Krakowie) stwierdzono liczne formy nanoplanktonu wapiennego wieku kampan-mastrycht. Wyznaczono poziomy nanoplanktonowe, określając wiek badanych osadów. W marglach bakulitowych, nazywanych też marglami z Węgierki, występujących w jednostce skolskiej, odnotowano podobny skład i charakter nanoplanktonu wapiennego jak w szarych marglach typu frydeckiego występujących w jednostce podśląskiej w zachodniej części Karpat. W badanych osadach jednostki podśląskiej nie stwierdzono ciągłego przejścia sedymentacyjnego między osadami mastrychtu i danu. Badania nanoplanktonu potwierdziły istnienie połączenia między strefą borealną i tetydzką na obszarze polskich Karpat zewnętrznych w kampanie i mastrychcie, o czym świadczy obecność w próbkach form nanoplanktonu uważanych za typowo borealne.
EN
In pelagic sediments of the Subsilesian Unit (Węglówka marls, Frydek-type marls, Żegocina marls and Rybie sandstones) and in the bakulit marls from the Skole Unit, and in marls from Bonarka in Silesian-Cracow Monocline many forms of calcareous nannoplankton of Campanian-Maastrichtian age were noted. Distinguishing of several nannoplankton assemblage zones allowed for more detailed time resolution of the studied sediments. Existing of the connection between the boreal and Tethyan realms in Polish Outher Carpathians during Campanian/Maastrichtian has been confirmed. Calcareous nannoplankton of the bakulit marls (Skole Unit) appeared similar to that from the Frydek-type marls (Subsilesian Unit). No continuous transition between sediments of the Maastrichtian and Danian within the studied Subsilesian Unit has been stated. Transition between Boreal and Tethyan realms on the Polish Outher Carpathians area, during Campanian and Maastrichtian age are confirmed.
EN
The heteromorph ammonite Nostoceras (Nostoceras) hyatti is described for the first time from the Piesting Formation (Gosau Group; Upper retaceous) at Grunbach along the eastern margin of the Northern Calcareous Alps, Lower Austria. This record significantly extends the geographic range of this late Campanian marker species; it was not previously known from the Alpine Cretaceous. Moreover, it corroborates earlier age assignments as late Campanian of the (lower) part of the Piesting Formation. A matrix sample taken from the specimen studied has yielded taxa whose range covers zones CC20 to CC23, or UC 15[^tp] to UC18, of the standard nannofossil zonation.
EN
The Campanian strata in the Wolbrom-Miechów area at Wierzchowisko, Jeżówka and Rzeżuśnia (i.e., the southwestern part of the Miechów Trough, southern Poland) have been studied in some detail. Collections of macrofossils available to date include generally well-preserved and diverse cephalopods (ammonoids, coleoids), inoceramid bivalves and irregular echinoids of considerable stratigraphic and correlative value. For the present paper, taxa which allow a preliminary correlation with northern Germany (Lagerdorf, Lehrte West Syncline and Munsterland Basin) are singled out for brief discussion. Stratigraphically useful taxa include the ammonites Pachydiscus (P.) haldemsis (SCHLUTER), Lewyites elegans (MOBERG), Scaphites (S.) hippocrepis III sensu COBBAN, S. (S.) gibbus SCHLUTER and Trachyscaphites spiniger spiniger (SCHLUTER), the coleoids Belemnitella ex gr. mucronata (VON SCHLOTHEIM) and Gonioteuthis quadrata (DE BLAINVILLE),the inoceramids Cataceramus dariensis (DOBROV & PAVLOVA), "Inoceramus" azerbaydjanensis ALIEV and "I." agdjakendsis ALIEV, and the echinoids Offaster pilula (LAMARCK), Galeola papillosa (LESKE), Echinocorys ex gr. subglobosa/turrita, E. ex gr. conica, Micraster (Gibbaster) ex gr. fastigatus/stolleyi and M. (M.) ex gr. schroederi/glyphus. The ammonite fauna, which is dominated by pachydiscids and diplomoceratids, is closely comparable to that from the Busko Zdrój area (i.e., the southeastern part of the Miechów Trough), but hoplitoplacenticeratids are still unknown from the Wolbrom-Miechów area which, taken together with inoceramid data, may point to a gap in the upper Lower Campanian (equivalent of conica/mucronata Zone).
EN
The nostoceratid ammonites Nostoceras (Didymoceras) postremum (BŁASZKIEWICZ, 1980) and Nostoceras (Didymoceras) varium (BŁASZKIEWICZ, 1980) are described from northern Germany for the first time. They appear with the second occurrence of Nostoceras (Bostrychoceras) polyplocum (ROEMER, 1841) in the upper portion of the Neancyloceras bipunctatum / Galerites roemeri Zone of the Lehrte West Syncline (Lower Saxony, east of Hannover), which is equivalent to the upper Belemnitella langei Zone of the chalk of Lńgerdorf-Kronsmoor (c. 50 km north of Hamburg). In northern Germany, this ammonite association appears c. 1 Ma earlier than Belemnella lanceolata, which marks the base of the Maastrichtian in belemnite terms, and c. 1.5 Ma earlier than the international base according to the first occurrences of Diplomoceras cylindraceum and Pachydiscus neubergicus at Lagerdorf-Kronsmoor.Assuming that the GSSP of the base of the Maastrichtian Stage at Tercis (southern France) equals the boundary of the U.S. Western Interior (radiometric age 71.3 . 0.5 Ma), Nostoceras (Bostrychoceras) polyplocum first appears in Europe c. 75 Ma before present.
EN
The revised geological log and the biostratigraphy of the Late Campanian and earliest Maastrichtian succession of the Middle Vistula River section, central Poland, are presented. The biostratigraphy is based on inoceramid fauna, basically newly collected. Eight inoceramid zones were distinguished, corresponding, in ammonite/belemnite terms, to an interval from the upper Bostrychoceras polyplocum Zone up to the traditional Belemnella occidentalis Zone. From bottom upwards these are zones of: Cataceramus subcompressus, "Inoceramus" tenuilineatus, Sphaeroceramus pertenuiformis, "Inoceramus" altus, "Inoceramus" inkermanaensis, Trochoceramus costaecus, "Inoceramus" redbirdensis, and of Endocostea typica. The inoceramid assemblages allow a precise correlation with critical sections in Europe, as well as trans-Atlantic correlation to the US Western Interior. The traditional base of the Maastrichtian falls at the base of the Trochoceramus costaecus Zone, whereas the 'Tercis basal Maastrichtian boundary' may fall as high as lower Belemnella sumensis Zone. Based on the correlation to the US Western Interior, a subdivision of the Upper Campanian into Middle and Upper substages is suggested. The boundary between the Middle and the Upper Campanian substages may be defined by the base of the "Inoceramus" tenuilineatus Zone, which in ammonite terms, corresponds to the base of the Didymoceras donezianum Zone (=top of the Bostrychoceras polyplocum Zone). Thirty one inoceramid species level taxa are described, of which "Inoceramus" smirnovi nom.nov. is a replacement name; 10 species are left in open nomenclature. Inoceramids of the Middle Vistula succession represent a uniform fauna characteristic of the whole Euramerican biogeographical region.
15
EN
Sixteen ammonite taxa are recorded from four temporary exposures of lower Upper Campanian deposits in the town of Busko Zdrój, NE limb of the Nida Trough, southern Poland. These are: Phylloceras (Neophylloceras) cf. bodei, Tetragonites obscurus, Desmophyllites sp., Pachydiscus (Pachydiscus) subrobustus, P. (P.) cf. subrobustus, Hoplitoplacenticeras (Hoplitoplacenticeras) dolbergense, H. (H.) sp., H. (Lemfoerdiceras) lemfoerdense, Glyptoxoceras cf. retrorsum, G. sp., Lewyites elegans, Pseudoxybeloceras (Pseudoxybeloceras) riosi, Ps. (Ps.) sp. juv., Baculites sp., Scaphites gibbus, and Trachyscaphites spiniger spiniger. Many specimens lack precise provenance data, but co-occurrence of T. obscurus, P. (P.) subrobustus, H. (H.) dolbergense, H. (L.) lemfoerdense, Ps. (Ps.) riosi, Baculites sp., S. gibbus, and T. s. spiniger is documented from a single opoka bed in one of the exposures.The ammonites allow correlation with standard sections in northern Germany: the ammonite-bearing sequence of Busko Zdrój corresponds to a part of the interval from the basiplana / spiniger to roemeri zones in Lńgerdorf and Kronsmoor (Schleswig-Holstein), and from the stobaei / basiplana to vulgaris / stolleyi zones in the Lehrte West Syncline (Lower Saxony). It can also be correlated with the lower part of the Neancyloceras phaleratum Zone in Vistula valley, central Poland.
EN
Late Cretaceous and Early Palaeogene echinoid faunules collected in recent years from the surface of an active mudflow at the Gschliefgraben near Gmunden (east of the Traunsee, Upper Austria) are both fairly diverse and of considerable palaeobiogeographic interest. So far, only (Late) Campanian taxa have been described and illustrated. In the present paper, notes are presented on additional Campanian, Maastrichtian and Palaeogene taxa, namely Lampadocorys? estermanni sp.nov., Lampadocorys? sp.nov. 1, Lampadocorys? sp.nov. 2, Rispolia cf. subtrigonata (CATULLO), Seunaster cf. heberti (SEUNES), Echinocorys ancileformis MOSKVIN & SHIMANSKAYA, Echinocorys ex gr. fonticola ARNAUD, Ganbirretia? sp., Micraster aturicus HEBERT in SEUNES, Micraster corcolumbarium DESOR, Micraster stolleyi (LAMBERT in DE GROSSOUVRE), Pseudogibbaster? sp., and Coraster beneharnicus SEUNES.These species indicate a close relationship with the Tethyan areas of northern Spain and SW France, the Crimea, Georgia and the northern Caucasus, although a number of holasteroid and micrasterid taxa would appear to be more common and widely distributed in boreal settings (e.g., NW Europe).
PL
Trzy odsłonięcia górnego kampanu, należące do trzech kolejnych poziomów amonitowych (Neancyloceras phaleratum, Bostrychoceras polyplocum i Didymoceras donezianum) w dolinie środkowej Wisły, zostały zbadane pod kątem paleośrodowiskowym. Bogata makrofauna ze wszystkich szczebli troficznych, świadczy o spokojnym środowisku, odpowiadającemu dzisiejszej strefie pogranicza wewnętrznego i zewnętrznego szelfu, o normalnym zasoleniu, dobrze natlenionych wodach przydennych i podłożu o zróżnicowanej konsystencji, stwarzającego różnorodnej faunie odmienne nisze ekologiczne.
EN
The paleoecology of macrofauna from three Upper Campanian outcrops in the Middle Vistula valley, belonging to three successive ammonite zones (Neancyloceras phaleratum, Bostrychoceras polyplocum and Didymoceras donezianum), have been investigated on paleoenvironmental ground. The rich macrofauna of all trophic levels indicates the calm epicratonic basin, corresponding to the inner to outer shelf environment, of normal salinity waters, good bottom-water oxygenation and with substrate of varied consistency, providing different niches for diverse fauna.
EN
Thirty-three inoceramid species from the upper Upper Campanian and Lower Maastrichtian of the Tercis section (SW France), the Global Boundary Stratotype Section and Point for the Campanian - Maastrichtian boundary, are described, of which 8 are left in open nomenclature. Two species are described as new: Endocostea jolkicevi and "Inoceramus" cobbani. The described species represent four inoceramid genera Cataceramus HEINZ, 1932, Sphaeroceramus, HEINZ, 1932, Endocostea WHITFIELD, 1877, Trochoceramus HEINZ, 1932, and thirteen species are referred to the genus "Inoceramus" sensu lato. The inoceramids allow the subdivision of the upper Upper Campanian and of the Lower Maastrichtian into seven zones and their correlation with the ammonite scheme, as well as with the chronostratigraphic standard. The Campanian - Maastrichtian boundary falls in the tompost part of the "Inoceramus" redbirdensis Zone, and the base of successive, Endocostea typica Zone is a good proxy for this stage boundary. Besides the Trochoceramus costaecus Zone all other zones are also distinguishable in the US Western Interior.
19
Content available remote Further Archaeogastropoda from the Campanian of Torallola, northern Spain
EN
18 archaeogastropod species are described from the Campanian of Torallola in north-eastern Spain, including 13 new species and one new genus. They are assigned to the Liotiinae, Colloniinae, Turbininae, Skeneidae and Cirridae. Our species of the Liotiinae are comparable with Recent as well as Triassic liotiids, the documented colloniids show close relations to moder species. Certain members of the Angaria-Astraea-Turbo-relation are difficult if not impossible to distinguish on shell characters alone, although these shells can be traced back into the Triassic. Three skeneiform species are assigned to Recent Skenea, but neither fossil nor recent Skoneidae are sufficiently well known, so our treatment remains preliminary. The new genus Torallochus is introduced for species having initially planispiral coiling that later changes to turriform coiling, and which are sculptured with strong ribs or spines on the periphery. The systematic position of this genus within the Trochidea remains uncertain. Among the Cirridae, the genusShikamancirrus in synonymized with the older name Sensuitrochus, and the youngest member of this genus is described. The new species are Pseudoliotina stinnesbecki, Arene mcleani, Homalopoma schroederi, Astraea iredalei,Astraea batalleria, Astraea hickmanae, Astraea? sohli, Skenea Wareni, Skenea suturata, Skenea torallensis, Torallochus rempensis, Torallochus pupiformis, and Torallochus discus.
20
Content available remote Trochidae (Archaeogastropoda) from the Campanian of Torallola in northern Spain
EN
30 species of the archaeogastropod family Trochidae are described from the Campanian of the southern Pyrenees, 15 of them are new. The new genus Amphigibbula is introduced and the genus Chilodontoidea is renamed as Hudledonta. Apparently several trochid lineages with living species can be traced back into the Late Cretaceous. This regards the Eucyclinae, Margaritinae, and Solariellinae, and the Tegulinae with some reservations. Among the groups examined here, only the members of the Trochinae appear to be of rather different character than their modern representatives. The fossil record of the Umboniinae can go far beyond that of the Late Cretaceous and connect even to Palaeozoic genera. The 15 new species are: Eucyclomphalus reminiscencius, Calliotropis torallolensis, Calliotropis seguris, Ilerdus pyrenaeus, Eucycloscala cretacea, Hudledonta nicolae, Danilia kosslerae, Margarites kasei, Margarites nielseni, Margarites kowalkei, Tectus quinteroi, Thoristella marshalli, Suavotrochus ponsi, Ethalia vinxae, and Protorotella herberti.
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