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EN
Multidisciplinary studies of the Middle-Upper Bathonian ore-bearing clays at Gnaszyn revealed variable palaeoenvironmental conditions during the deposition of this seemingly monotonous sequence. We interpret the conditions in the bottom environment and the photic zone, and also evaluate the influence of the adjacent land areas, based on sedimentology, geochemistry, sporomorphs and palynofacies composition, benthic (foraminifera, gastropods, bivalves, scaphopods, echinoderms), planktonic (calcareous nannoplankton, dinoflagellate cysts), and nektonic (sharks) fossils. The Gnaszyn succession originated relatively close to the shore, within reach of an intense supply of terrestrial fine clastic and organic particles. The latter are mainly of terrestrial origin and range from 1.5 to 2.5 wt.%. The precise water depth is difficult to estimate but most likely ranges from several tens of metres to a few hundred metres. All fossil groups show minor changes throughout the succession. As the climate seems to have been quite stable during this period we consider sea-level fluctuations to have been the main factor responsible for the changes. The terrestrial input, including freshwater and land-derived clastic and organic particles (sporomorphs and cuticles), increased during periods of sea-level lowstand. As a consequence, stress conditions (lower salinity, higher nutrient availability, lower water transparency) in the photic zone caused blooms of opportunistic planktonic taxa. Furthermore, a faster sedimentation rate led to oxygen depletion and deterioration of the living conditions in the bottom environment due to an increased accumulation of organic matter. As a result, the benthic biota became taxonomically impoverished and commonly dominated by juvenile forms. During periods of high sea level, the source areas were shifted away from the basin, resulting in a decrease in the terrestrial influx, increase in the salinity of surface waters, the appearance of more diverse phytoplankton assemblages, a lower sedimentation rate, and an improvement of living conditions at the bottom.
EN
Systematic sampling through the Middle and Upper Bathonian strata at Gnaszyn has resulted in the discovery of 13 neoselachian teeth. Systematically, the teeth represent five taxa including Sphenodus sp., Protospinax sp. 1, Protospinax sp. 2, Palaeobrachaelurus sp. and another, indeterminate orectolobiform. The presence of two species of the flattened and bottom-dwelling Protospinax and two different orectolobiforms that are likely to have lived near the bottom, is a strong indication of oxygenated bottom conditions at the time of deposition. The dietary preferences of these taxa included a wide variety of benthic invertebrates. The synechodontiform Sphenodus may have been the first pelagic predatory neoselachian in the Jurassic, equipped with high and slender piercing teeth that formed a tearing-type dentition. The diet of Sphenodus probably included bony fish, smaller sharks and cephalopods.
EN
As many hybodont sharks are known solely from their teeth, this investigation approaches the phylogeny of the group with an emphasis on tooth morphology and dentitional patterns. The preliminary results presented here suggest that at least four different lineages of hybodont sharks occurred in the Mesozoic. Dentitional characters imply a close relationship within the Lonchidiidae (Lonchidion, Vectiselachos, Parvodus, and tentatively Hylaeobatis), within the Hybodontinae (Hybodus and Egertonodus) and in another, unnamed subfamily of the Hybodontidae, including Planohybodus, Secarodus and Priohybodus. There is also weak support for a grouping of Acrodus, Asteracanthus and Palaeobates in the Acrodontinae, while Lissodus is left without family designation due to a rather unique dentition and cephalic spine morphology. "Polyacrodus" is considered a nomen dubium as there are no characters to diagnose the genus based on the type species.
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