The Order Clupeiformes Bleeker, 1859 comprises herrings, anchovies, sprats, sardines, and shads. The fossil record of this group is rich within the Paratethys. Here we describe a new clupeiform fish, †Sanalosa janulosa gen. et sp. nov., from the Lower Oligocene of the Carpathian Basin, Poland. This new genus has a unique combination of characters (lower jaw articulation located under the posterior part of the orbit; abdominal scutes well developed with 3 to 5 in the gular region, 11–14 prepelvic scutes associated with ribs, 11–12 postpelvic scutes; several striae on the frontals; an opercle with 6–12 thin radial ridges; a horizontal ramus of the preopercle shorter than the vertical one; 42–44 vertebrae; 8–10 supraneurals; a dorsal fin with 18–22 rays, and an anal fin with 21–23 rays), supporting recognition of a new genus and species within the Family Alosidae Svetovidov, 1952. Similarities and differences between fossil and extant genera of the Clupeiformes are discussed to shed more light on their relationship. Moreover, the palaeobiogeography, diversity and distribution of Oligocene clupeiform fishes in the Paratethys are presented and discussed.
The Suborder Clupeoidei Bleeker, 1859 comprises mostly marine fishes, which form large schools feeding on plankton. The fossil record of the suborder in the Late Paleogene reveals that clupeoids were abundant in the Western, Central, as well as Eastern Paratethys. Clupeoid descriptions from the Polish Outer Carpathians (SE Poland, Central Paratethys) remain incomplete due to usage of ‘collective species’, taxonomical inaccuracies, and lack of clear links between extinct and extant representatives. In this paper we present a new clupeoid, †Beksinskiella gen. nov., from the Oligocene of the Outer Carpathians, Poland. The new genus encompasses the nominal species †Maicopiella longimana (Heckel, 1850). †Beksinskiella gen. nov. has a unique combination of characters (skull roof with frontoparietal striae; smooth opercle; 6–7 branchiostegal rays; 42–48 vertebrae; dorsal and anal fins with 21–23 rays; the last two rays of the anal fin being elongated, and the presence of a complete series of abdominal scutes with the postpelvic ones poorly developed), supporting recognition of a new genus of the Suborder Clupeoidei. Similarities and differences between fossil and extant genera of Cluepoidei are discussed to shed light on their relationship.
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