Pinus cembra forests are limited to the Alps and Carpathians. Although several studies regarding their structure were carried out in some locations in the Tatra Mts. it required further investigations. Therefore, the aims of this study were to describe the stand and shrub structure of P. cembra forests, compare their structure with the Picea abies forests and analyse differences between silicate and calcicolous P. cembra forests in the Tatra Mts. The data were collected on the 16 sampling plots (500 m2), in the Swiss stone pine and Norway spruce forests. We measured the diameter at breast height (dbh) of each tree and recorded the young trees and shrubs. In order to compare species composition between silicate and calcicolous P. cembra forests, we made 91 relevés in their entire range of distribution (917 ha). Furthermore, we examined the share of main tree species along the altitude and inclination gradients, using the GAM models. The tree density in the P. cembra forests reaches 618 stems per ha, whereas their basal area (BA) 23.17 m2 ha-1. Main tree species are P. cembra and P. abies. P. cembra dominates in the higher thickness classes. The BA and dbh structure varies significantly between P. cembra and P. abies forests. The most abundant juveniles are P. abies and Sorbus aucuparia. The differences between forests growing on different substrate are relatively low. The altitude has a significant impact on the share of P. cembra (increase) and P. abies (decrease). The inclination has a significant impact on the increase of share of P. cembra.
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The frequently changed temperature could have great effects on soil fauna community during soil thawing period in cold areas. Therefore, soil faunas were investigated in both the soil organic layer (OL) and mineral soil layer (ML) in the primary fir (Abies faxoniana) forest (PF), fir and birch (Betula albosinensis) mixed forest (MF) and secondary fir forest (SF) in the eastern Qinghai–Tibet Plateau every ten days between March 5 and April 25, 2009. Soil macrofauna was picked up by hand in the fields. Mesofauna was collected and separated from the soil samples by Baermann and Tullgren methods, respectively. The dominated species of macrofauna belonged to Coleoptera and Diptera at the early stage of soil thawing, and to Coleoptera, Diptera, Araneae and Hymenoptera at the later stage. However, the dominated species of mesofauna belonged to Nematode, Collembola, and Acari in the whole soil thawing. The density, number of taxa, and diversity index of soil fauna showed significant change with temperature fluctuations and reached an obvious peak when soil temperatures rising above 0[degrees]C. The density and number of taxa of macrofauna in both the OL and ML were the lowest on March 5 in the three forests, but the density of mesofauna in both the OL and ML was the highest on March 25, except for the ML in the PF. These results implied that soil fauna community was sensitive to temperature fluctuations, which is important in understanding the ecological processes in the winter–spring transitional period.
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Natural regeneration of forest depends on the light regimes of floor. Point-based methods such as fisheye photo and radiometer can not provide a full panorama of light regime of heterogeneous forest stand. Eastern Tibetan Plateau is a major forest belt characteristic of diverse forest type and topographic differentiation. Understanding the trend of changes of light regime along succession series of forest may be helpful for the management of ecosystems. Fragmented forest patches due to tectonic activity and human intervention have made this prediction difficult. We use a spatially explicit forest stand light model (tRAYci) to simulate light distribution within forest in typical subalpine forest succession series of eastern Tibetan Plateau. Due to the spatial heterogeneity of tree distribution in the subalpine area, the forest stand can be approximated with a spatially explicit model of trees. Three typical subalpine forest stands (Sabina forest (SF), Fir forest (FF) and Birch forest (BF)) are selected in the eastern Tibetan Plateau. The dominant species are sabina (Sabina saltuaria (Rehd. et Wils.) Cheng), fir (Abies faxoniana Rehd. et Wils.) and birch (Betula platyphylla Suk.) for each stand and they are spatially clumped in distribution. They represent old growth coniferous forest (SF, 330 years old), coniferous-broadleaved forest (FF, 180 ys) and pioneer broadleaved forest (BF, 40 ys). The parameters of the three-dimensional model of trees are calibrated with field measurements. The simulated values are generally consistent with observed values of radiation measured by radiometers installed in these stands and values derived from fisheye photos. Test failures may be caused by the incomplete submodel of crown as a gap free one. Light regimes in old growth and pioneer forest are much more heterogeneous than intermediate stages of forest. Light regimes of these forests are also reflected by the composition of understory herb layers.
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