For the first time, a quantitative analysis of the land-plant diversity during deposition of the Rotliegend (latest Pennsylvanian-Early Permian; Gzhelian - Sakmarian) of the Saar-Nahe basin (SW-Germany) and the Zechstein (Late Permian; Wuchiapingian) of the German Zechstein basin is given. A study of six taphofloras indicates that regionally not only the overall species richness declined, but also the diversity as represented by individual taphofloras. This decline occurred after the deposition of the Upper Rotliegend (Asselian-Sakmarian), leading to an extremely low species richness during the deposition of the Zechstein. Besides taphonomic differences between the indi- vidual localities considered here, possible reasons for the observed lower diversity include most probably a climatic deterioration/aridisation due to tectonic northwards movement of Pangaea. As compared to the Late Pennsylvanian and the Early Permian, this climatic deterioration together with a contemporaneous decrease in topographic relief due to erosion and collapse of the Variscan Orogen, may also be the main reason for a decrease in habitat diversity and finally the decrease in land-plant diversity. However, based on the data presented here it is difficult to judge whether the observed regional diversity decline at individual localities can be generalized for the entire Euramerican floral province. Nevertheless, it can be concluded from other sources, that there are large gaps in the palaeobotanical record of this region, which may lead to underestimation of the "true" land plant diversity during this period within the Euramerican floral province.
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