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1

Taxonomic problems of some zonal species of the genus Palmatolepis and the correlation of the Frasnian of the East European Platform with conodont zonations

Ovnatanova Nonna S., Kononova Lyudmila I.

Acta Geologica Polonica

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2020

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Vol. 70, no. 1

107--124

EN

Most of the Frasnian regional stages of the Ukhta region, South Timan, Russia, are composed of basinal deposits, however, the hassi and jamieae zones of the Standard Conodont Zonation cannot be easily recognised in this region. A revision of the previously elaborated succession of the Timan-Pechora associations revealed that the correlation problems are connected with the taxonomic interpretation of the zonal species Palmatolepis hassi and Palmatolepis jamieae. Analysis of the conodont collection of Professor W. Ziegler from the Rhenish Slate Mountains, Germany, especially from the interval encompassing the Lower hassi to Upper rhenana zones, has provided evidence of the lack of validity of the jamieae Zone as a separate stratigraphic unit. This statement is based on the composition of the jamieae and Lower rhenana conodont associations and the absence of P. jamieae near the lower boundary of the zone in the stratotype and other sections of the Rhenish Slate Mountains, becoming more common upsection. The correlation between the Timan-Pechora conodont associations III–XI, the Standard Conodont Zonation (Ziegler and Sandberg 1990) and the Frasnian Zonation (Klapper 1989; Klapper and Kirchgasser 2016) is suggested herein. The Domanikian Regional Stage corresponds to the punctata–Late hassi zones of the Standard Conodont Zonation and to Frasnian Zones 5–10. The boundaries of Frasnian Zones 8–9 need to be further specified in South Timan. The correlation between the Standard Conodont Zonation and the Frasnian Zonation of Klapper is elaborated.

2

Brachiopods and stratigraphy of the Upper Devonian (Frasnian) succession of the Radlin Syncline (Holy Cross Mountains, Poland)

Baliński A., Racki G., Halamski A. T.

Acta Geologica Polonica

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2016

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Vol. 66, no. 2

125--174

EN

The lower part of the Frasnian succession in the Radlin Syncline (Kielce–Łagów Synclinorium, southern region of the Holy Cross Mountains), in the two studied successions: Józefka at Górno and (for the first time) Radlin, consists of the rhythmic marly Szydłówek Beds, the fossil-rich limestones of the Wietrznia Beds (locally) and the atypically developed, calcareous Kostomłoty Beds. The carbon isotope chemostratigraphic pattern overall corresponds well to the global Early–Middle Frasnian biogeochemical perturbation, even if the major punctata positive excursion is only fragmentarily recorded in the Kostomłoty intrashelf basin. Two brachiopod assemblages are abundantly represented in both sections: the Phlogoiderhynchus polonicus Assemblage, typical of the Szydłówek Beds, and the Biernatella lentiformis Assemblage, limited to the middle part of the Wietrznia Beds. Both are highly dominated by the index species. Twenty nine lower Frasnian brachiopod species (Craniida – 1 species, Strophomenida – 1, Productida – 2, Protorthida – 1, Orthida – 5, Pentamerida – 1, Rhynchonellida – 4, Atrypida – 4, Athyridida – 3, Spiriferida – 4, Spiriferinida – 3) are described from the Szydłówek and Wietrznia Beds. Seven new species are introduced: Skenidioides cretus Halamski sp. nov., Biernatium minus Baliński sp. nov., Monelasmina montisjosephi Baliński sp. nov., Atryparia (Costatrypa) agricolae Halamski and Baliński sp. nov., Davidsonia enmerkaris Halamski sp. nov., Leptathyris gornensis Baliński sp. nov., and Echinocoelia parva Baliński sp. nov. Davidsonia enmerkaris Halamski sp. nov. is intermediate between Davidsonia Bouchard-Chantereaux, 1849 and Rugodavidsonia Copper, 1996 and is the youngest known representative of the suborder Davidsonioidea Copper, 1996. Skenidioides cretus Halamski sp. nov. is the last representative of the genus. Statistical investigation of a large sample of Spinatrypina (Exatrypa) explanata did not confirm the existence of two dimorphic forms, coarse- and fine-ribbed. The high-diversity Biernatella lentiformis Assemblage is quite dissimilar to coeval brachiopod assemblages described heretofore from the Holy Cross Mountains region. It is interpreted as consisting of mostly parautochthonous dwellers of deep-slope muddy habitats and a local, occasionally storm-agitated, intra-basin brachiopod-crinoid-coral shoal. The fauna was adapted probably to cooler and nutrient-poor waters during an initial phase of the severe carbon cycle perturbation.

3

Upper Devonian microvertebrates from the Canning Basin, Western Australia

Roelofs B., Playton T., Barham M., Trinajstic K.

Acta Geologica Polonica

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2015

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Vol. 65, no. 1

69--100

EN

A diverse microvertebrate fauna is described from the Virgin Hills and Napier formations, Bugle Gap Limestone Canning Basin, Western Australia. Measured sections at Horse Spring and Casey Falls (Virgin Hills Formation) and South Oscar Range (Napier Formation) comprise proximal to distal slope carbonates ranging in age from the Late Devonian Frasnian to middle Famennian. A total of 18 chondrichthyan taxa are identified based on teeth, including the first record of Thrinacodus tranquillus, Cladoides wildungensis, Protacrodus serra and Lissodus lusavorichi from the Canning Basin. A new species, Diademodus dominicus sp. nov. is also described and provides the first record of this genus outside of Laurussia. In addition, the upper range of Australolepis seddoni has been extended to Late Devonian conodont Zone 11, making it the youngest known occurrence for this species. The Virgin Hills and Napier formations microvertebrate faunas show close affinities to faunas recovered from other areas of Gondwana, including eastern Australia, Iran, Morocco and South China, which is consistent with known conodont and trilobite faunas of the same age.

4

Pelagic chondrichthyan microremains from the Upper Devonian of the Kale Sardar section, eastern Iran

Hairapetian V., Ginter M.

Acta Geologica Polonica

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2010

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Vol. 60, no. 3

357-371

EN

An assemblage of fifteen taxa of chondrichthyan microremains from late Frasnian through late Famennian pelagic deposits of the Kale Sardar section, eastern Iran, is described. Several taxa (Phoebodus bifurcatus, Phoebodus sophiae and Protacrodus vetustus) are reported for the first time from Iran. The presence of Deihim mansureae and Ph. sophiae in the Late rhenana to linguiformis Zones and Phoebodus rayi in the early triangularis Zone of the Kale Sardar section, provides new biostratigraphic information. The late Frasnian part of the assemblage corresponds to the faunas from the intrashelf basins of central europe, and the middle.late Famennian part is comparable to that from the tafilalt Platform of Morocco.

5

Septacja w muszlach ślimaków frańskich z Gór Świętokrzyskich

Krawczyński W.

Przegląd Geologiczny

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2009

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Vol. 57, nr 1

39-39

EN

Amongst several known Paleozoic gastropod species having transverse septa in the apical parts of shells, three species (Straparollus aff. circularis, Orecopia kadzielniae and Donaldiella karczewskii) have been found in the Frasnian limestones of Grabina and Bolechowice-Panek in the Holy Cross Mountains (Central Poland). Septation in gastropod shells is a rare phenomenon and is an adaptive feature. Septa occur either in the thick-shelled gastropods living in high-energetic (reef) environments, preventing against negative effects of destruction of the apical parts of shells, or in the thin-shelled gastropods living on muddy substrate, lowering the inner volume of a habitable part of shell and increasing outer surface of gastropod body.

6

Charakterystyka sedymentologiczna górnej części warstw szydłóweckich

Vierek A.

Przegląd Geologiczny

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2008

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Vol. 56, nr 9

848-856

EN

The Givetian to Frasnian boundary interval in the N part of the Kielce region (Holy Cross Mts.) consists of dark-coloured shales, marls and micritic limestones defined as Szydłówek Beds. The upper parts of the Szydłówek Beds are well exposed in the Kostomłoty-Mogiłki and Górno-Józefka quarries. These sections are situated in Kostomłoty transitional facies zone between the shallow-water carbonate platform and the deeper Łysogóry basin. The uppermost parts of the Szydłówek Beds comprise many fine-grained intercalations: laminated calcisiltites, calcarenites, calcirudites and coquinas beds. The shales, marls and micritic limestones are hemipelagic deposits of deep, quiet and oxygen-depleted basin, whereas grained limestones correspond with shallower water and higher energy environment. Laminated calcisiltites and finer calcarenites originated in the environment below storm wave base and are interpreted as surge-like turbidity flows deposits. Thicker calcarenites, calcirudites, and coquinas beds were deposited about storm wave base and are interpreted as tempestites.

7

The mid-Frasnian subsidence pulse in the Lublin Basin (SE Poland) : sedimentary record, conodont biostratigraphy and regional significance

Narkiewicz K., Narkiewicz M.

Acta Geologica Polonica

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2008

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Vol. 58, no. 3

287-301

EN

Most of the thickness of Frasnian sediments in the central segment of the Lublin Basin, i.e. up to 300 metres, is represented by a single transgressive-regressive Cycle VIa, developed in the carbonate-sulphate platform facies. The age of the transgressive part falls into the interval between the upper part of theUpper hassi Zone and the jamieae Zone, whereas the upper boundary runs between the upper part of the Lower rhenana Zone and the lower part of the Upper rhenana Zone. Basin architecture and conodont biostratigraphic data confirm the tectonic nature of the cycle, which represents a short-termincrease in Frasnian subsidence and depositional rates. Based on the conodont data, it is plausible that the onset of the tectonic subsidence in the Lublin Basin and the incipient Pripyat Graben rifting correspond closely in age. They can thus be attributed to the common tectonic mechanism of regional extension in the south-west part of the East European Platform. The lack of any Late Devonian magmatic activity in the Lublin Basin and the synchronous development of this basin with the Pripyat Graben favour the idea that intraplate stresses were the primary factors controlling subsidence in both depocentres during the mid Frasnian to Famennian. The hypothetical mantle plume could have merely amplified the effects of crustal extension in the Pripyat Graben, thus facilitating a typical rift development.

8

Early Frasnian sharks from central Iran

Hairapetian V., Ginter M., Yazdi M.

Acta Geologica Polonica

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2008

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Vol. 58, no. 2

173-179

EN

Two limestone bone-beds in the early Frasnian of the Chahriseh section, central Iran, yielded numerous chondrichthyan teeth and scales. The fauna includes, most probably, only two taxa: a hitherto unknown aztecodontid omalodontiform, Manberodus fortis gen. et sp. nov., and a multicuspid phoebodontiform, provisionally referred to as Phoebodus cf. latus GINTER & IVANOV, 1995.Anew omalodontiform family, Aztecodontidae, including Aztecodus LONG& YOUNG, 1995 and Manberodus gen. nov., is proposed.

9

Devonian filter-feeding sharks

Ginter M.

Acta Geologica Polonica

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2008

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Vol. 58, no. 2

147-153

EN

Upper Frasnian rocks of Utah and Nevada yielded several multicuspid, low-crowned shark teeth. It is proposed that they were used mainly for filtering food from water. Two new chondrichthyan species bearing such teeth were distinguished: a phoebodontid Diademodus utahensis sp. nov., with up to 17 very delicate cusps in the tooth-crown; and a cladodont of uncertain systematic position, Lesnilomia sandbergi gen. et sp. nov., also known from the upper Frasnian of Moravia.

10

Zapis zmian w zespołach małżoraczkowych w osadach granicy fran-famen w Górach Świętokrzyskich

Olempska E.

Przegląd Geologiczny

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2003

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Vol. 51, nr 11

948-986

PL

: W Górach Świętokrzyskich badania nad zmianami zespołów małżoraczkowych na granicy fran-famen prowadzone były w facjach umiarkowanie głębokiego basenu śródszelfowego w profilu Płucki (Olempska 2002) oraz płytszych środowiskach w profilu Kostomłoty (Casier i in., 2000) i profilu Psie Górki w Kielcach (Casier i in., 2002). Gatunki planktonowych małżoraczków z rodziny Entomozoidae występują licznie w późnofrańskich i wczesnofameńskich wapienno-marglistych osadach w profilu Płucki. Zmiany w zespole małżoraczkowym na granicy fran-famen w tym profilu wykazują wyraźne zubożenie zespołu w tzw. interwale kryzysowym i następują w dwu etapach. Część gatunków wymiera u podstawy wapienia głowonogowego (Górny Horyzont Kellwasser), natomiast główny etap wymierania następuje w trakcie sedymentacji wapienia głowonogowego. Pojawienie się w wapieniu głowonogowym gatunku Entomoprimitia (Entomoprimitia) kayseri,'egzotycznego' dla tego obszaru, wiąże się z krótkotrwałym pulsem transgresywnym mającym miejsce w poziomie linguiformis. Główny puls wymierania entomozoidów wiąże się prawdopodobnie z krótkotrwałym pulsem regresywnym. Entomozoidy powracają na ten obszar jako nowe gatunki z obszarów refugiów z początkiem pulsu transgresywnego w środkowym poziomie triangularis. Zespół fameński zdominowany jest przez masowo wystepujacy gatunek Franklinella (Franklinella) sigmoidale. Badania prowadzone nad zmianami zespołów bentosowych małżoraczków w profilach Kostomłoty (Casier i in., 2000) i Psie Górki (Casier i in., 2002) wykazały, że około 75% gatunków małżoraczków bentosowych wymarło na granicy fran-famen w rejonie kostomłockim i ponad 70% w profilu Psich Górek. Zdaniem badaczy tych sekwencji, decydującym czynnikiem powodującym wymieranie fauny małżoraczków na tych obszarach było ochłodzenie oraz zmiany poziomu morza.

EN

The entomozoacean ostracod species are common in late Frasnian to early Famennian rhythmic calcareous-marly sequence in the Plucki section, Holy Cross Mts., Poland. The entomozoacean faunal change across the Frasnian-Famennian boundary shows the overall reduction in species number in two steps in the latest Frasnian. A few species became extinct at the base of the Upper Kellwasser Horizon, and the main extinction was within the UKWH. The appearance of E. (E.) kayseri in the UKWH coincides with a short-term transgressive pulse in the linguiformis Zone. The final extinction of entomozoaceans at the boundary, coincides with the short-term regressive pulse. The entomozoaceans reappear as new species from refugia lineages at the beginning of the transgressive pulse in the Middle triangularis Zone. The results of the Frasnian-Famennian extincion and recovery of benthic ostracods in the Kostomłoty and Psie Górki sections presented by Casier et al. (2000) and Casier et al. (2002) are briefly summarized.