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1

Life span and growth rate of Middle Jurassic mesohibolitid belemnites deduced from rostrum microincrements

Wierzbowski H.

Volumina Jurassica

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2013

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Vol. 11, no. 1

1--18

EN

The microincrements of nine Middle Jurassic mesohibolitid belemnite rostra (Hibolithes jurensis, Pachybelemnopsis fusiformis and Pachybelemnopsis “subhastatus”) were analysed to measure the ontogenic age and growth rate of the belemnites. The microincrements are interpreted to have been formed daily based on analogy with extant coleoid cephalopods and the frequency of circadian rhythms in animal activity. The mesohibolitid belemnites are found to have had a short life span of ca. 1 year. The short life span of the belemnites studied is similar to the life span of modern coleoids (squids, cuttlefishes and common octopuses). It restricts the utility of belemnite rostra as a proxy for average secular seawater temperatures and chemistry. Short-lived cycles (2–8 days) in the belemnite growth rate are linked to variations in the metabolic activity of these animals. Minor diagenetic alteration revealed by the dull cathodoluminescence of some rostra is found to have no effect on the preservation of the growth rings.

2

The Belemnella stratigraphy of the Campanian-Maastrichtian boundary; a new methodological and taxonomic approach

Remin Z.

Acta Geologica Polonica

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2012

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Vol. 62, no. 4

495-533

EN

The taxonomy and stratigraphy of the Upper Campanian and Lower Maastrichtian belemnites from the Vistula (central Poland) and Kronsmoor (northern Germany) sections are revised on the basis of new collections from the Vistula section as well as a reinvestigation of the classic collection of Schulz from the Kronsmoor section. For the taxonomic description a new biometric procedure is proposed, which can be applied to both the genera Belemnella and Belemnitella. For the species-level taxa recognition the Artificial Neural Networks method, the self-organizing Kohonen algorithm, was implemented. This new taxonomic and methodological approach enabled the recognition of nine species of the genus Belemnella. Five of them can be assigned to the existing species B. lanceolata, B. longissima, B. inflata, B. obtusa and B. vistulensis. However, the species concept differs from that applied by Schulz (1979). As a consequence, the stratigraphic ranges of these species are modified. Four species are left in open nomenclature and represent possibly new species. Future studies may reveal that they might be assigned to East European forms from Ukraine or Russia. The species of Belemnella recognized are placed into the stratigraphic framework based on the standard ammonite and inoceramid bivalve zonations, especially those recognized in the Vistula section. The newly proposed belemnite zonation for the Vistula and Kronsmoor sections is correlated via inoceramids with the standard GSSP at Tercis, France, in order to identify the base of the Maastrichtian Stage. The Campanian/Maastrichtian boundary as defined in Tercis is placed here at the base of the newly defined B. obtusa and B. vistulensis Zones ["obtusa/vistulensis"] - thus it is markedly higher than the traditional boundary based on the FAD of representatives of the genus Belemnella - This new boundary coincides well with a distinct turnover of belemnite guard morphology and represents one of the most important points in the early evolutionary history of Belemnella. Three belemnite zones defined by their lower boundaries are recognized in the Campanian/Maastrichtian interval, in addition to three subzones recognized within the B. obtusa Superzone. The B. lanceolata and B. inflata zones as understood here are referred to the Upper Campanian [Tercis definition]. The B. obtusa Zone is subdivided into three subzones, viz.: Belemenlla vistulensis, Belemnella sp. G and Belemnella sp. F, which are referred to the Lower Maastrichtian [Tercis definition]. The fast evolving species of Belemnella enable the proposal of a biostratigraphic scheme with a resolution that is higher than those based on inoceramid bivalves and ammonites - the longevity of a belemnite zone could be as low as 200Ky.

3

Integrated stratigraphy of the Kronsmoor section (northern Germany), a reference point for the base of the Maastrichtian in the Boreal Realm

Niebuhr B., Hampton M., Gallagher L., Remin Z.

Acta Geologica Polonica

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2011

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Vol. 61, no. 2

193-214

EN

Results of detailed multistratigraphic analyses of the Campanian.Maastrichtian boundary section at Kronsmoor in northern Germany are summarised and calibrated with the GSSP at Tercis les Bains, southwest France. Additional markers for the definition of the boundary in the Boreal Realm are proposed, and a detailed carbon isotope curve around the Campanian.Maastrichtian boundary in the chalk facies of the Boreal epicontinental sea is presented. The C isotopic GSSP marker for global correlation is the markedly abrupt decrease of c. 0.7 [per mil] [delta^13]C directly at the Campanian.Maastrichtian boundary as dated by ammonites. In electronic borehole measurements the Kronsmoor section covers the SP peaks 53 to 64 and the base of the Maastrichtian being situated just below SP peak 60. The first occurrence (FO) of the ammonite Pachydiscus neubergicus, which corresponds to biohorizon 1 at Tercis, falls in the upper part of nannofossil Zone UC15, at both localities. Biohorizon 3 is the FO of the ammonite Diplomoceras cylindraceum, which first appears in the Upper Campanian of Tercis and at Kronsmoor enters significantly above the FO of Belemnella lanceolata, the conventional Boreal belemnite marker for the base of the Maastrichtian Stage. Based on ammonite evidence, the internationally accepted base of the Maastrichtian at Kronsmoor is located between the FOs of Diplomoceras cylindraceum (Upper Campanian) and Pachydiscus neubergicus (Lower Maastrichtian) c. 11 m above flint layer F 600, at which the first representatives of the belemnite genus Belemnella, in particular Bn. lanceolata occur. The latter thus is a Late Campanian species, appearing c. 450 ky prior to the ammonite-based boundary. The FOs of Belemnella pseudobtusa (sensu Schulz) resp. Belemnella obtusa (sensu Remin) directly at the boundary can be use as the coleoid proxy for the definition of the base of the Maastrichtian in the Boreal Realm. To define the boundary by benthic foraminifera the last occurrence (LO) of Neoflabellina praereticulata is suitable. Biohorizon 12, as defined at Tercis, involves the nannofossil Uniplanarius trifidus, however, at Kronsmoor this species is rare, occurs only sporadically and also significantly lower in comparison to Tercis. It is possible though to compare and correlate nannofossil events between Kronsmoor and Tercis using cosmopolitan taxa such as Broinsonia parca constricta and Eiffellithus eximius. The LO of the latter appears to be situated just above the boundary in both sections; it follows from this that the top of nannofossil Zone UC15 is of Early Maastrichtian age.

4

Occurrence and significance of Cenomanian belemnites in the lower Danubian Cretaceous Group (Bavaria, southern Germany)

Wilmsen M., Niebuhr B., Chellouche P.

Acta Geologica Polonica

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2010

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Vol. 60, no. 2

231-241

EN

The belemnite records of the lower Danubian Cretaceous Group (DCG, northeastern Bavaria, southern Germany) are compiled, taxonomically described and placed within the new integrated stratigraphic framework of the group. Three specimens from the lower Regensburg Formation (Saal Member) south of Regensburg can be assigned to Neohibolites cf. ultimus (d'Orbigny) and are dated as late Early Cenomanian (Mantelliceras dixoni Zone). Eight specimens represent Praeactinocamax plenus (Blainville) and occur in an event (plenus Event) in the lower Eibrunn Formation (Regensburg area) or basal Regensburg Formation (Roding area in the Bodenwohrer Senke). Biostratigraphy and carbon stable isotopes suggest that the belemnite horizon with P. plenus in the DCG has strictly the same chronostratigraphic position (mid-Late Cenomanian, middle Metoicoceras geslinianum Zone) as elsewhere in Central and NW Europe. The lithostratigraphic units of the lower Danubian Cretaceous Group (i.e., the Regensburg and Ebirunn formations), however, are characterized by a pronounced diachronism based on their time-transgressive (i.e., onlapping) deposition during the Cenomanian.Early Turonian transgression. The distribution of P. plenus around the Mid-European Island can be easily explained by migration around the positive area without the necessity of a marine strait across the Bohemian Massif.

5

Sztuczne sieci neuronowe Kohonena jako narzędzie w taksonomii paleontologicznej - metodyka oraz zastosowanie na przykładzie późnokredowych belemnitów

Remin Z.

Przegląd Geologiczny

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2008

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Vol. 56, nr 1

58-66

EN

Artificial neural networks (ANNs), the computer software or systems that are able to "learn" on the basis of previously collected input data sets are proposed here as a new useful tool in paleontological modeling. Initially ANNs were designed to imitate the structure and function of natural neural systems such as the human brain. They are commonly used in many natural researches such as physics, geophysics, chemistry, biology, applied ecology etc. Special emphasis is put on the Kohonen self-organizing mapping algorithm, used in unsupervised networks for ordination purposes. The application of ANNs for paleontology is exemplified by study of Late Cretaceous belemnites. The Kohonen networks objectively subdivided the belemnite material] ~ 750 specimens) into consistent groups that could be treated as monospecific. The possibility of transferring these results to the language of classical statistics is also presented. Further development and possibility of use of ANNs in various areas of paleontology, paleobiology and paleoecology is briefly discussed.