Palynostratigraphic studies of the Carboniferous strata of the Szczawno and Wałbrzych formations, outcropping in the vicinity of Wałbrzych in the northern part of the Intra-Sudetic Basin, are reported. Rocks of the Szczawno Formation in a section near Konradów had earlier been assigned to the Upper Visean, while rocks of the Wałbrzych Formation in the Biały Kamień section were included to the Lower Namurian [the Stenozonotriletes triangulus-Rotaspora knoxi (TK) Miospore Zone]. This study describes abundant palynological material with many taxonomically diverse miospore assemblages from these rocks, allowing precise palynostratigraphic conclusions. These showed that the rocks of the Szczawno Formation studied should be referred to the lower part of the Serpukhovian [Verrucosisporites morulatus (Vm) Subzone], correlated with the lowest Namurian, while rocks of the Wałbrzych Formation from Biały Kamień belong to the Lower Bashkirian [Lycospora subtriquetra-Cirratriradites rarus (SR) Subzone], correlated with the Upper Namurian A. These results indicate that the rocks from both sections studied are younger than they were earlier believed. This revises the age of the Szczawno and Wałbrzych formations in the northern part of the Intra-Sudetic Basin which belong to four miospore zones, from Vm to SR, corresponding to the Serpukhovian and lowermost Bashkirian (Namurian A corresponding to the Pendleian to Alportian of British chronostratigraphy).
The known occurrence of corals distinguished here in the new Family Krynkaphyllidae varies at the subfamily level. Those of the Subfamily Krynkaphyllinae subfam. nov. are so far almost unknown from outside of the Donets Basin. In contrast, those of the Subfamily Colligophyllinae subfam. nov. are common, possibly ranging from the lower Viséan Dorlodotia Salée, 1920, a potential ancestor of the family, to the Artinskian Lytvophyllum tschernovi Soshkina, 1925. They bear different generic names, but were all originally described as fasciculate colonial. A detailed study of Lytvophyllum dobroljubovae Vassilyuk, 1960, the type species of Colligophyllum gen. nov., challenges that recognition in that at least some of those taxa are solitary and gregarious and/or protocolonial. As such, solitary, protocolonial and, probably, fasciculate colonial habits are accepted in the Colligophyllinae subfam. nov., whereas the Krynkaphyllinae subfam. nov. contains only solitary taxa. The resemblance to the Suborder Lonsdaleiina Spasskiy, 1974 led to the analysis of families included in that suborder by Hill (1981) in the context of their relationship, or homeomorphy, to Krynkaphyllidae fam. nov. This question primarily concerns the Family Petalaxidae Fomichev, 1953; a relationship with the Family Geyerophyllidae Minato, 1955, is more distant, if one exists. The distinct, parallel stratigraphic successions of taxa within two subfamilies of the Krynkaphyllidae fam. nov. document their probably common roots and early divergence. However, a lack of robust data precludes an interpretation or treatment of those successions as phylogenetic. The absence of key stratigraphic and morphologic data meant that eastern Asiatic taxa have not been considered in these successions; however, morphological similarities allow for their tentative inclusion within the Krynkaphyllidae fam. nov. The following new taxa are introduced: Krynkaphyllidae fam. nov., Krynkaphyllinae subfam. nov., Colligophyllinae subfam. nov., Krynkaphyllum gen. nov., Colligophyllum gen. nov., Protokionophyllum feninoense sp. nov., Krynkaphyllum multiplexum sp. nov., Krynkaphyllum validum sp. nov., and three species of Protokionophyllum Vassilyuk in Aizenverg et al., 1983 left in open nomenclature.
Seven genera (one new), belonging to four subfamilies, seven named species (six new), four species left in open nomenclature and two specimens included in this paper as unnamed Aulophyllidae are described from strata ranging from the lowermost Bashkirian Limestone D510 to the lower Bashkirian Limestone F1. A new genus: Voragoaxum and six new species: Dibunophyllum medium, Dibunophylloides columnatus, D. paulus, D. similis, Voragoaxum cavum and Rozkowskia lenta are introduced. Comparison of the ontogeny of the earliest Bashkirian species of Nina Fedorowski, 2017a and Dibunophylloides Fomichev, 1953 suggest the derivation by descent of the Family Bothrophyllidae from the Subfamily Dibunophyllinae. This means that true bothrophylla are absent in the Mississippian strata of the Western European Province and, perhaps, in the contemporaneous strata of other areas as well.
Cordibia pumila gen. et sp. nov. (Aulophyllidae, Dibunophyllinae) from the Lower Bashkirian E1 Limestone of the Donets Basin, Ukraine is described. The term "protocolony" is formally defined as a growth form intermediate between the solitary and colonial. Protocolonies consist of protocorallites and verticils of non-reproductive offsets [i.e., the lost structures]. The early ontogeny of a protocorallite is compared to the blastogeny in the same species in order to establish characters in common during both processes and to document the usefulness of blastogeny for phylogenetic reconstructions. The short duration and abundance of the occurrence (Limestones E1up acme, E11) of that species also makes it a good marker for the lowest Feninian (= Krasnopolyanian) strata in the Donets Basin.
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Two successions of Carboniferous sedimentary rocks in Central Iran yielded ammonoid assemblages from one horizon each. In the Ramsheh section, a siltstone with Neoglyphioceras yazdii n. sp. and Dombarites can be assigned to the latest Visean. The assemblage from the Shesh Angosht Mountain is more diverse with species of the genera Proshumardites, Eumorphoceras, Homoceras, Glaphyrites, Syngastrioceras, and Dombarocanites. This is the first time that the co-occurrence of the genera Eumorphoceras and Homoceras has been recorded. This fauna contains elements of the South Urals and Central Asian occurrences of Bashkirian ammonoids and suggests close palaeobiogeographic relationships.
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