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The Middle Oxfordian to lowermost Kimmeridgian ammonite succession at Mikhalenino (Kostroma District) of the Russian Platform, and its stratigraphical and palaeobiogeographicai importance

Głowniak E., Kiselev D., Rogov M., Wierzbowski A., Wright J.

Volumina Jurassica

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2010

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Vol. 8, no. 1

5-48

EN

The Mikhalenino section on the Russian Platform has yielded numerous ammonites from the Middle and Upper Oxfordian and lowermost Kimmeridgian, collected bed by bed. The ammonites belong mostly to the Boreal family Cardioceratidae, but also to the Subboreal family Aulacostephanidae; additionally at some levels there were collected various Submediterranean ammonites (Perisphinctidae, Oppeliidae and Aspidoceratidae). The co-occurrence of ammonites representative of different faunal provinces makes possible recognition of the Boreal, Subboreal, and partly also Submediterranean standard zonations. In consequence, it is possible to make a close correlation between these zonal schemes. The Oxfordian/Kimmeridgian boundary at the Pseudocordata/Baylei zonal boundary of the Subboreal zonal scheme corresponds precisely to the Rosenkrantzi/Bauhini zonal boundary. This boundary of the stages defined well faunistically in the Flodigarry section (Isle of Skye, Scotland) and proposed as a candidate for the uniform Oxfordian/Kimmeridgian boundary, can be also recognized in the Russian section studied. The boundary can be traced in the Mikhalenino section using the same criteria as used at Staffin: the appearance of the first representatives of Pictonia [M]–Prorasenia [m] (Subboreal), and the first appearance of Amoeboceras (Plasmatites) (Boreal). This indicates the large correlation potential of the boundary defined in this way. The research on the Mikhalenino section has provided the new palaeontological findings described in this study. These include the first discovery in the territory of the Russian Platform of ammonites of the genera/subgenera Decipia, Vineta and Pomerania (Pachypictonia). Two new species: Decipia (?) kostromensis sp. nov., and Pictonia mesezhnikovi n. sp. are described.

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The ammonite faunas of the Callovian-Oxfordian boundary interval in Europe and their relevance to the establishment of an Oxfordian GSSP

Page K., Melendez G., Wright J.

Volumina Jurassica

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2009

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Vol. 7, no. 1

89-99

EN

At Moenkopi Wash along the Ward Terrace escarpment of northern Arizona strata of the upper Dinosaur Canyon Member of the Moenave Formation contain sedimentary structures we interpret as casts of tetrapod burrows. Sandstone casts and in situ burrows occur concentrated in two horizons that extend several hundred meters along the Ward Terrace escarpment. The structures, hosted in beds of eolian sandstone, form interconnecting networks of burrows that branch at right angles. Individual burrow casts have sub-circular cross sections and consist of nearvertical tunnels and horizontal to low-angle galleries that connect to larger chambers. Most burrow casts measure 5 to 15 cm in diameter, are filled by sandstone of similar grain size as the host rock, and have walls that are unlined and lack external ornamentation. Bedding plane exposure of the lower horizon reveals that the density of burrows exceeds 30 vertical tunnels per square meter. One exposure in the upper horizon reveals burrows concentrated in a mound-like structure with 1 m of relief. Rhizoliths, distinguished from burrows by their typical smaller diameters, calcareous infilling, and downward branching, co-occur with these burrows in the upper horizon. The fossil burrows in the Moenave Formation appear to have been constructed by a fossorial tetrapod with social behavior similar to the modern Mediterranean blind mole-rat. Although no skeletal remains are associated with the burrows, the fossil record suggests that the most likely producers of the Moenave burrows were tritylodontid cynodonts.

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Integrated stratigraphical study of the candidate Oxfordian Global Stratotype Section and Point (GSSP) at Redcliff Point, Weymouth, Dorset, UK

Page K., Melendez G., Hart M., Price G., Wright J., Bown P., Bello J.

Volumina Jurassica

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2009

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Vol. 7, no. 1

101-111

EN

Ham Cliff near Redcliff Point, Weymouth, Dorset (SW England) exposes one of Europe's most complete Callovian-Oxfordian boundary sequences and has consequently been identified as a potential candidate GSSP for the base of the Oxfordian Stage. The boundary sequence lies within the thick mudrock facies of the Oxford Clay Formation and is abundantly fossiliferous, cardioceratid ammonites in particular being conspicuous. By convention, the stage boundary is drawn at the first occurrence of the genus Cardioceras here represented by C. redcliffense Page, Melendez and Wright at the base of the Scarburgense Subchronozone of the Mariae Chronozone. Associated Perisphinctoidea (including Peltoceras, Alligaticeras and Euaspidoceras) provide additional biostratigraphical information. Other macrofossil groups show less discernible changes, although frequent belemnites (Hibolithes) provide new highresolution carbon and strontium isotope data which are consistent with global curves and continuous sedimentation across the boundary interval. Magnetostratigraphic information is also available. Foraminiferal assemblages are dominated by epistominids but include a flood of early planktonic forms, including ?Globuligerina oxfordiana (Grigelis) immediately above the boundary. Well-preserved nannofloras are dominated by Watznaueria with conspicuous Zeugrhabdotus, podorhabdids and Stephanolithion indicating the NJ14 Biozone. Ostracoda and holothurian spicules are also recorded. These results are synthesised to provide a multidisciplinary, integrated review of the suitability of Redcliff Point for the definition of an Oxfordian GSSP. Correlations with the French candidate site in Haute-Provence are discussed and proposals made for formally establishing a GSSP for the base of the Oxfordian Stage in Europe.

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The ammonite faunas of the Callovian/Oxfordian boundary interval in Europe and their relevance to the establishment of an Oxfordian GSSP

Melendez G., Page K., Wright J., Atrops F.

Volumina Jurassica

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2006

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Vol. 4, no. 1

185-185

EN

The latest Callovian and Early Oxfordian represent one of the most dynamic intervals in the history of Jurassic Ammonoidea and is characterized by one of the highest levels of mixing of Boreal, Submediterranean and even Mediterranean faunas. In particular the massive expansion of Boreal Cardioceratidae from their original “home” in Arctic areas as far south as South East France, brings them into contact with Mediterranean-style faunas rich in Phylloceratidae. This so-called “Boreal Spread” (after J. H. Callomon) provides the framework within which high-resolution inter-bioprovincial correlations are possible and hence the context for a sucessful GSSP designation for the base of the Oxfordian Stage within Europe (and hence the beginning of the Upper Jurassic). Associated with the Cardioceratidae, however, is a great variety of Perisphinctina, including Aspidoceratidae, Periphinctidae, Grossouvridae and rarer Pachyceratidae as well as frequent Hecticoceratidae and rarer Phylloceratidae. The latter groups are much more abundant in southern areas (Tethyan Realm), but the Aspidoceratidae do persist well into the Boreal Realm. Crucially, several groups of the Perisphinctina persist beyond Europe and therefore provide tantalising indications that a truly global correlation of any GSSP established in Europe will ultimately be possible. The current paper will review the stratigraphical, taxonomic and palaeobiogeographical context and significance of the trans Callovian/Oxfordian boundary faunas within Europe, building on recent results from the UK and France. Conclusions will be drawn concerning the appropriate – or convenient – level at which the place the Callovian-Oxfordian in Europe and its potential interpretation elsewhere. Such conclusions are highly relevant to the eventual establishment of an Oxfordian GSSP.

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Integrated stratigraphical study of the candidate Oxfordian Global Stratotype Section and Point (GSSP) at Redcliff Point, Weymouth, Dorset, UK

Page K., Melendez G., Hart M., Price G., Wright J., Bown P., Bello J.

Volumina Jurassica

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2006

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Vol. 4, no. 1

200-201

EN

Redcliff Point near Weymouth, Dorset (SW England) exposes one of Europe’s most complete Callovian-Oxfordian boundary sequences and has been the subject of a rigorous multidisciplinary assessment. The boundary sequence lies entirely within the clay facies of the Oxford Clay Formation, the relatively high carbonate content of which facilitates the excellent preservation of both macro- and microfaunas (and floras) as well as geochemical information. Ammonites, in particular, are conspicuous, and partly retain an aragonitic shell. By convention, the stage boundary is drawn at the first occurrence of the genus Cardioceras, which has been interpreted as corresponding to the transition between “Quenstedtoceras” paucicostatum (Lang) and Cardioceras ex gr. scarburgense (Young and Bird), specifically at the first occurrence of C. woodhamense Arkell sensu Callomon (non Marchand). This transition is well seen at Redcliff and provides the primary means through which the boundary can be correlated. Associated Perisphinctina (including Peltoceras, Alligaticeras, Properisphinctes and Euaspidoceras) provide additional biostratigraphical information. Other macrofossil groups show less discernible changes, although the end of the Callovian in England marks the local, virtual disappearance of Boreal cylindroteuthid belemnites with the persistence of Tethyan hibolithids into the Early Oxfordian. Isotopic studies of recovered belemnites record important information on carbon and strontium isotopes and provide new, high resolution data for the refinement of the global curves. The isotope data are also consistent with continuous sedimentation across the boundary. Foraminiferal assemblages are dominated by poorly preserved epistominids. Planktonic Foraminifera are recorded, mainly as pyrite steinkerns. This makes identification difficult although a flood close to the boundary appears to be Globuligerina oxfordiana. Other planktonic taxa are present, including one species that may be new. Nannofloras are well preserved, common to abundant and dominated by Watznaueria britannica with conspicuous Zeugrhabdotus erectus, podorhabdids and Stephanolithion bigotii. The presence of Stephanolithion bigotii maximum throughout, places the samples within the NJ14 biozone. Ostrocoda and holothurian spicules are also recorded. These results are synthesised to provide a multidisciplinary, integrated review of the suitability of Redcliff Point for the definition of an Oxfordian GSSP. Correlations with the French candidate site in Savournon, Haute-Provence are discussed and proposals made for formally establishing a GSSP for the base of the Oxfordian Stage in Europe.

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A potential stratotype for the Oxfordian/Kimmeridgian boundary: Staffin Bay, Isle of Skye, UK

Wierzbowski A., Coe A., Hounslow M., Matyja B., Ogg J., Page K., Wierzbowski H., Wright J.

Volumina Jurassica

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2006

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Vol. 4, no. 1

17-33

EN

A coastal exposure of the Staffin Shale Formation at Flodigarry, Staffin Bay, Isle of Skye, Scotland, UK fulfils the criteria for definition as the Global Stratotype Section and Point (GSSP) for the base of the Kimmeridgian Stage (Upper Jurassic). This marine shale succession was deposited during a long-term transgression, and is part of a complete, relatively well-expanded stratigraphic succession. A rich fauna of ammonites above and below the Oxfordian/Kimmeridgian boundary allows recognition of the Evoluta Subzone (Pseudocordata Zone) and Rosenkrantzi Subzone (Rosenkrantzi Zone) of the Subboreal and Boreal uppermost Oxfordian, and the Densicostata Subzone (Baylei Zone) and the Bauhini Zone of the Subboreal and Boreal lowermost Kimmeridgian). A suitable level for the boundary is thus marked by the replacement of the Subboreal Ringsteadia (M)/Microbiplices (m) by Pictonia (M)/Prorasenia (m), and by the first appearance of Boreal Amoeboceras (Plasmatites). Detailed study of the microfossils reveals an excellent dinoflagellate succession. A variety of stratigraphically important dinoflagellates are found, the assemblages being intermediate in character between Boreal and Subboreal ones. The magnetostratigraphic data, though rather troublesome to extract, shows a polarity pattern which can be confidently correlated to other UK boundary sections. The upper boundary of a normal magnetozone falls at, or very near, the proposed Oxfordian/Kimmeridgian boundary. The 87Sr/86Sr ratio at the boundary, based on an analysis of belemnites, lies between 0,70689 and 0,70697, averaging 0.70693. Matching worldwide trends, no distinct change in the ratio is seen across the boundary. A lack of variations in the carbon isotope composition of belemnites across the Oxfordian/Kimmeridgian boundary does not indicate perturbation in the global carbon cycle. However, high ?13C values and their scatter suggest the influence of local fractionation affecting isotope composition of dissolved inorganic carbon (DIC) in the partly isolated Boreal sea. A fall in the belemnite ?18O values in the Upper Oxfordian and Lower Kimmeridgian compared to the Mid Oxfordian suggests a slight rise in seawater temperature.