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Osteology of a large allosauroid theropod from the Upper Jurassic (Tithonian) Morrison Formation of Colorado, USA

Dalman S.

Volumina Jurassica

2014

Vol. 12, no. 2

159--180

Two partial skeletons of allosaurid theropods belonging to an adult and a juvenile from the Upper Jurassic (Tithonian) Morrison Formation of McElmo Canyon in Montezuma County, southwestern Colorado, were discovered in 1953 by the late Joseph T. Gregory and David Techter. The adult specimen consists of several isolated cranial and postcranial skeletal elements that are exceptionally well-preserved and include the left premaxilla, maxilla, dentary, teeth, quadratojugal, two caudal vertebrae, pubic peduncle, ischium, proximal tibia, a nearly complete left foot, and several isolated teeth, whereas the juvenile specimen is represented by the distal portion of the right dentary and a fragmentary splenial. The specimens represent a new species of Allosaurus, here named Allosaurus lucasi, which differs from Allosaurus fragilis by having a relatively short premaxilla and robust quadratojugal with short jugal process and a short quadrate process of the quadratojugal that is at the same level as the rostral quadratojugal ramus. The presence of a new species of Allosaurus in the Tithonian of North America provides further evidence of the taxonomic and morphological diversity of the Allosauridae clade and their continuous evolutionary success, which extended to the Cretaceous.

New data on small theropod dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA

Dalman S.

Volumina Jurassica

2014

Vol. 12, no. 2

181--196

In 1879, Othniel C. Marsh and Arthur Lakes collected in the Upper Jurassic Morrison Formation Quarry 12 at Como Bluff, Wyoming, USA, several isolated axial and appendicular skeletal elements of small theropod dinosaurs. Since the discovery the specimens remained unnoticed for over a century. The skeletal remains of small theropods are rare at Como Bluff and throughout the Morrison Formation. Their bones are delicately constructed, so they are not as well-preserved as the bones of large-bodied theropods. The bones of small theropods described here were found mixed with isolated crocodile teeth and turtle shells. Comparison of the skeletal materials with other known theropods from the Morrison Formation reveals that some of the bones belong to a very small juvenile Allosaurus fragilis and Torvosaurus tanneri and also to a new ceratosaur taxon, here named Fosterovenator churei, whereas the other bones represent previously unidentified juvenile taxa of basal tetanuran and coelurid theropods. The discovery and description of these fossil materials is significant because they provide important information about the Upper Jurassic terrestrial fauna of Quarry 12, Como Bluff, Wyoming. The presence of previously unidentified theropod taxa in the Morrison Formation indicates that the diversity of basal tetanuran and coelurid theropods may have been much greater than previously expected. Although the fossil material here described is largely fragmentary, it is tenable that theropods of different clades co-existed in the same ecosystems at the same time and most likely competed for the same food sources.