New data on small theropod dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA

• Autorzy: Dalman S.

Identyfikatory

DOI

10.5604/17313708 .1130142

• Języki publikacji: EN

Abstrakty

EN

In 1879, Othniel C. Marsh and Arthur Lakes collected in the Upper Jurassic Morrison Formation Quarry 12 at Como Bluff, Wyoming, USA, several isolated axial and appendicular skeletal elements of small theropod dinosaurs. Since the discovery the specimens remained unnoticed for over a century. The skeletal remains of small theropods are rare at Como Bluff and throughout the Morrison Formation. Their bones are delicately constructed, so they are not as well-preserved as the bones of large-bodied theropods. The bones of small theropods described here were found mixed with isolated crocodile teeth and turtle shells. Comparison of the skeletal materials with other known theropods from the Morrison Formation reveals that some of the bones belong to a very small juvenile Allosaurus fragilis and Torvosaurus tanneri and also to a new ceratosaur taxon, here named Fosterovenator churei, whereas the other bones represent previously unidentified juvenile taxa of basal tetanuran and coelurid theropods. The discovery and description of these fossil materials is significant because they provide important information about the Upper Jurassic terrestrial fauna of Quarry 12, Como Bluff, Wyoming. The presence of previously unidentified theropod taxa in the Morrison Formation indicates that the diversity of basal tetanuran and coelurid theropods may have been much greater than previously expected. Although the fossil material here described is largely fragmentary, it is tenable that theropods of different clades co-existed in the same ecosystems at the same time and most likely competed for the same food sources.

Słowa kluczowe

EN

dinosaurs; theropoda; Upper Jurassic; Morrison Formation; Como Bluff; western USA

PL

dinozaury; teropody; górna jura; Formacja Morrisona; Como Bluff; zachodnie USA

• Wydawca: Państwowy Instytut Geologiczny - Państwowy Instytut Badawczy
• Czasopismo: Volumina Jurassica
• Rocznik: 2014
• Tom: Vol. 12, no. 2
• Strony: 181--196
• Opis fizyczny: Bibliogr. 68 poz.

Twórcy

autor

Dalman S.

• Department of Geosciences, Fort Hays State University, 600 Park Street, Hays, KS 67601, USA

Bibliografia

• 1.AZUMA Y., CURRIE P.J., 2000 — A new carnosaur (Dinosauria: Theropoda) from the Lower Cretaceous of Japan. Canadian Journal of Earth Sciences, 37: 1735–1753.
• 2.BAKKER R.T., 1986 — The dinosaur heresies. William Morrow, New York.
• 3.BAKKER R.T., 1998 — Brontosaur killers: Late Jurassic allosaurids as sabre-tooth cat analogues. Gaia, 15: 145–158.
• 4.BAKKER R.T., KRALIS D., SIEGWARTH J., FILLA J., 1992 — Edmarka rex, a new, gigantic theropod dinosaur from the middle Morrison Formation, Late Jurassic of the Como Bluff outcrop region. Hunteria, 2: 1–24.
• 5.BENSON R.B.J., 2008 — New information of Stokesosaurus, a tyrannosauroid (Dinosauria: Theropoda) from North America and the United Kingdom. Journal of Vertebrate Paleontology, 28: 732–750.
• 6.BENSON R.B.J., 2009 — A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods. Zoological Journal of the Linnean Society, doi: 10.1111/j.1096-3642. 2009.00569.x.
• 7.BENSON R.B.J., 2010 — The osteology of Magnosaurus nethercombensis (Dinosauria, Theropoda) from the Bajocian (Middle Jurassic) of the United Kingdom and a re-examination of the oldest records of tetanurans. Journal of Systematic Palaeontology, 8: 131–146.
• 8.BENSON R.B.J., CARRANO M.T., BRUSATTE S.L., 2009 — A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic. Naturwissenschaften, 97: 71–78.
• 9.BRUSATTE S.L., BENSON R.B.J., 2013 — The systematic of Late Jurassic tyrannosauroid theropods from Europe and North America. Acta Palaeontologica Polonica, 58: 47–54.
• 10.BRUSATTE S.L., BENSON R.B.J., HUTT S., 2008 — The osteology of Neovenator salerii (Dinosauria: Theropoda) from the Wealden Group (Barremian) of the Isle of Wight. Palaeontographical Society Monograph, 162: 1–75.
• 11.BRITT B.B., 1991 — Theropods of Dry Mesa Quarry (Morrison Formation, Late Jurassic), Colorado, with emphasis on the osteology of Torvosaurus tanneri. Brigham Young University Geology Studies, 37: 1–72.
• 12.CARPENTER K., MILES C., CLOWARD K., 2005a — New small theropod from the Upper Jurassic Morrison Formation of Wyoming. In: The Carnivorous Dinosaurs (Ed. K. Carpeneter): 23– 48. Indiana University Press.
• 13.CARPENTER K., MILES C., OSTROM J.H., CLOWARD K., 2005b — Redescription of the small maniraptoran theropods Ornitholestes and Coelurus from the Upper Jurassic Morrison Formation of Wyoming. In: The Carnivorous Dinosaurs (Ed. K. Carpenter): 49–71. Indiana University Press.
• 14.CARRANO M.T., 2007 — The appendicular skeleton of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology Memoir, 8: 163–179.
• 15.CARRANO M.T., BENSON R.B.J., SAMPSON S.D., 2012 — The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology, 10: 211–300.
• 16.CARRANO M.T., SAMPSON S.D., 2008 — The phylogeny of Ceratosauria (Dinosauria: Theropoda). Journal of Systematic Palaeontology, 22: 510–534.
• 17.CHURE D.J., 1994 — Koparion douglassi, a new dinosaur from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument; the oldest troodontid (Theropoda: Maniraptora). Brigham Young University Geology Studies, 40: 11–15.
• 18.CHURE D.J., 1995 — A reassessment of the gigantic theropod Saurophagus maximus from the Morrison Formation (Upper Jurassic) of Oklahoma, USA. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers: 103–106.
• 19.CHURE D.J., 2001 — The second record of the African theropod Elaphrosaurus (Dinosauria, Ceratosauria) from the Western Hemisphere. Neues Jahrbuch für Geologie und Paläontologie Monatshefte, 9: 566–576.
• 20.CROSS C.W., 1894 — Pike’s Peak Colorado. Geological Atlas Folio 1894, United States Geological Survey, 8.
• 21.CURRIE P.J., ZHAO X.-J., 1994 — A new carnosaur (Dinosauria: Theropoda) from the Jurassic of Xinjiang, People’s Republic of China. Canadian Journal of Earth Sciences, 30: 2037–2081.
• 22.CURRIE P.J., AZUMA Y., 2006 — New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan. Journal of Paleontological Society of Korea, 22: 173–193.
• 23.DALMAN S.G., 2014 — Osteology of a large allosauroid theropod from the Upper Jurassic (Tithonian) Morrison Formation of Colorado, USA. Volumina Jurassica, 12, 2: 159–180 (this volume).
• 24.DODSON P., BEHRENSMEYER A.K., BAKKER R.T., MCINTOSH J.S., 1980 — Taphonomy and paleoecology of the dinosaur beds of the Jurassic Morrison Formation. Paleobiology, 6: 208–232.
• 25.FOSTER J., 2003 — Paleoecological analysis of the vertebrate fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain Region, USA. New Mexico Museum of Natural History and Science Bulletin, 23: 1–95.
• 26.FOSTER J., 2007 — The gargantuan to minuscule: the Morrison menagerie, part II. In: Jurassic West, the dinosaurs of the Morrison Formation and their world (Ed. J.O. Farlow). Indiana University Press: 162–242.
• 27.FOSTER J.R., CHURE D.J., 2000 — An ilium of a juvenile Stokesosaurus (Dinosauria, Theropoda) from the Morrison Formation (Upper Jurassic: Kimmeridgian), Meade County, South Dakota. Brigham Young University Geology Studies, 45: 5–10.
• 28.FOSTER J.R., CHURE D.J., 2006 — Hindlimb allometry in the Late Jurassic theropod dinosaur Allosaurus, with comments on its abundance and distribution. New Mexico Museum of Natural History and Science Bulletin, 36: 119–122.
• 29.GALTON P.M., 1980 — Elaphrosaurus, an ornithomimid dinosaur from the Upper Jurassic of North America and Africa. Paläontologische Zetischrift, 56: 265–272.
• 30.GALTON P.M., 1982 ― Elaphrosaurus, an ornithomimid dinosaur from the Upper Jurassic of North America and Africa. Paläontologische Zeitschrift, 56: 265–275.
• 31.GALTON P.M., JENSEN J.A., 1979 — A new large theropod dinosaur from the Upper Jurassic of Colorado. Brigham Young University Geology Studies, 26: 1–12.
• 32.GAUTHIER J., 1986 — Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences, 8: 1–55.
• 33.HARTMAN S., LOVELACE D., WAHL D., 2005 — Phylogenetic assessment of a maniraptoran from the Morrison Formation. Journal of Vertebrate Paleontology, 25: 67A–68A.
• 34.HENDRICKX C., MATEUS O., 2014 — Torvosaurus gurneyi n. sp., the largest terrestrial predator from Europe, and a proposed terminology of the maxilla anatomy in nonavian theropods. PLoSONE, 9: 1–25.
• 35.HOLTZ T.R., JR., MOLNAR R.E., CURRIE P.J., 2004 — Basal Tetanurae. In: The Dinosauria. Second Edition (eds D.B. Weishampel et al.): 71–110. University of California Press, Berkeley.
• 36.HUENE F.V. von, 1914 — The dinosaurs not a natural order. American Journal of Science, Series 4 38: 145–146.
• 37.HUENE F.V. von, 1920 — Bemerkungen zur Systematik und Stammesgeschichte einiger Reptilien. Zeitschrift für Induktive Abstammungs- und Vererbungslehre, 24: 209–212.
• 38.HUNT A.P., LUCAS S.G., 2006 — A small theropod dinosaur from the Upper Jurassic of eastern New Mexico with a checklist of small theropods from the Morrison Formation of western North America. New Mexico Museum of Natural History and Science Bulletin, 36: 115–118.
• 39.HUTT S., 1999 — Neovenator salerii: a new theropod dinosaur from the Wealden of the Isle of Wight: its status and significance for theropod evolution [MS Thesis]. University of Portsmouth, Portsmouth.
• 40.HUTT S., MARTILL D.M., BARKER M.J., 1996 — The first European allosaurid dinosaur (Lower Cretaceous, Wealden Group, England). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 1996: 635–644.
• 41.JENSEN J.A., 1981 — Another look at Archaeopteryx as the world’s oldest bird. Encyclia, 58: 109–128.
• 42.JENSEN J.A., 1985 — Uncompahgre dinosaur fauna: A preliminary report. Great Basin Naturalist, 45: 710–720.
• 43.JENSEN J.A., PADIAN K., 1989 — Small pterosaurs and dinosaurs from the Uncompahgre fauna (Brushy Basin Member, Morrison Formation, ?Tithonian), Late Jurassic, Western Colorado. Journal of Paleontology, 63: 364–373.
• 44.KOWALLIS B.J., CHRISTIANSEN E.H., DEINO A.L., PETERSON F., TURNER C.E., KUNK M.J., OBRADOVICH J.D., 1998 — The age of the Morrison Formation. Modern Geology, 22: 235–260.
• 45.LITWIN R.J., TURNER C.E., PETERSON F., 1998 — Palynological evidence on the age of the Morrison Formation, Western Interior U.S. Modern Geology, 22: 297–319.
• 46.LOEWEN M.A., 2004 — Variation and stratigraphic distribution of Allosaurus within the Late Jurassic Morrison Formation. Geological Society of America Abstracts with Programs, 36: 524A.
• 47.LOEWEN M.A., 2009 — Variation in the Late Jurassic theropod dinosaur Allosaurus: Ontogenetic, functional, and taxonomic implications [PhD Thesis]. University of Utah.
• 48.MADSEN J.H. Jr., 1974 — A new theropod dinosaur from the Upper Jurassic of Utah. Journal of Paleontology, 48: 27–31.
• 49.MADSEN J.H. Jr, 1976 — A second new theropod dinosaur from the Late Jurassic of east central Utah. Utah Geological and Mineral Survey, Utah Geology, 3: 51–60.
• 50.MADSEN J.H. Jr, 1993 — Allosaurus fragilis a revised osteology. Utah Geological and Mineral Survey Bulletin, 109: 1–163.
• 51.MADSEN J.H., JR., WELLES S.P., 2000 — Ceratosaurus (Dinosauria, Theropoda) a revised osteology. Utah Geological Survey, Miscellaneous Publication, 00-2: 1–80.
• 52.MALAFAIA E., DANTAS P., ORTEGA F., ESCASO F., 2007 — Nuevos restos de Allosaurus fragilis (Theropoda: Carnosauria) del yacimiento de Andrés (Jurásico Superior; centro-oeste de Portugal). In: Cantera Paleontológica: 255–271. Diputación Provincal de Cuenca, Cuenca.
• 53.MARSH O.C., 1877 — Notice of some new dinosaurian reptiles from the Jurassic Formation. American Journal of Science, Series 3, 14: 514–516.
• 54.MARSH O.C., 1878 — Principal characters of American Jurassic dinosaurs ― part 1. American Journal of Science, Series 3, 16: 411–416.
• 55.MARSH O.C., 1879 — Notice of new Jurassic reptiles. American Journal of Science, Series 3, 18: 501–505.
• 56.MARSH O.C., 1881 — Principal characters of American Jurassic dinosaurs ― part 5. American Journal of Science, Series 3, 21: 417–423.
• 57.MARSH O.C., 1884 — Principal characters of American Jurassic dinosaurs ― part 8; the Order Theropoda. American Journal of Science, Series 3, 27: 329–340.
• 58.MATEUS O., ANTUNES M.T., 2000 — Torvosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. Livro de Resumos de I Congreso Ibérico de Pleontologia/XVI Jornadas de la Sociedad Española de Paleontologia: 115–117.
• 59.MATEUS O., WALEN A., ANTUNES M.T., 2006 — The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to that of the Morrison Formation, with a description of a new species of Allosaurus. New Mexico Museum of Natural History and Science Bulletin, 36: 123–129.
• 60.OSBORN H.F., 1903 — Ornitholestes hermanii, a new compsognathoid dinosaur from the Upper Jurassic. Bulletin of the American Museum of Natural History, 19: 459–464.
• 61.OSTROM J.H., 1980 — Coelurus and Ornitholestes: are they the same? In: Aspects of Vertebrate History (Ed. L. Jacobs): 245–256. Flagstaff, Museum of Northern Arizona Press.
• 62.PÉREZ-MORENO B.P., CHURE D.J., PIRES C., MARQUES DA SILVA C., PÓVOAS L., CACHÃO M., SANZ J.L., GALOPIM de CARVALHO A.M., 1999 — On the presence of Allosaurus fragilis (Theropoda: Carnosauria) in the Upper Jurassic of Portugal: first evidence of an intercontinental dinosaur species. Journal of the Geological Society, London, 156: 449–452.
• 63.RAUHUT O.W.M., 2003 — The interrelationships and evolution of basal theropod dinosaurs. The Palaeontological Association, Special Papers in Palaeontology, 69: 1–213.
• 64.RAUHUT O.W.M., 2005 — Post-cranial remains of ‘coelurosaurs’ (Dinosauria, Theropoda) from the Late Jurassic of Tanzania. Geological Magazine, 142: 97–107.
• 65.RAUHUT O.W.M., FECHNER R., 2005 — Early development of the facial region in a non-avian theropod dinosaur. Proceedings of the Royal Society of London, Series B, 272: 1179–1183.
• 66.SEELEY H.G., 1887 — On the classification of the fossil animals commonly named Dinosauria. Proceedings of the Royal Society of London, 43: 165–171.
• 67.SIEGWARTH J.D., LINDBECK R.A., REDMAN P.D., SOUTHWELL E.H., BAKKER R.T., 1997 — Giant carnivorous dinosaurs of the family Megalosauridae from the Late Jurassic Morrison Formation of eastern Wyoming. Contributions from the Tate Museum Collections, Casper, Wyoming, 2: 1–33.
• 68.TURNER C.E., PETERSON F., 1999 ― Biostratigraphy in the Upper Jurassic Morrison Formation of the Western Interior, U.S.A. In: Vertebrate Paleontology in Utah (Ed. D.D. Gillette): 77– 114. Utah Geological Survey Miscellaneous Publication 99-1.