First nearly complete skeleton of the Cretaceous duvaliid belemnite Conobelus

• Autorzy: Lukeneder A.
• Języki publikacji: EN

Abstrakty

EN

The first Cretaceous belemnite preserved with the rostrum, slightly compressed phragmocone and part of the proostracum is described from the Early Cretaceous (Late Valanginian) Rossfeld Formation (Eibeck, Reichraming Nappe of the Northern Calcareous Alps). The rostrum has dorsal groove (alveolar furrow) typical of duvaliids, and its conical shape (round in transverse section outline), and the rounded apex allow its attribution to Conobelus Stolley, 1919. The new species C. pseudoheres sp. nov. is proposed based on the unique features of the specimen, i.e. persistently parallel lateral sides throughout the rostrum and the conical, blunt outline at the apex. The ration between rostrum and phragmocone is 0.78; the distances between the calcitic septa of the anterior end of the phragmocone range from 3 to 5 mm. The proostracum is 0.02 mm thick. The apical angle is 32° and the alveolar angle (posterior end of the phragmocone) is 24°.The alveolus is 40 mm long, yielding a ratio between rostrum and alveolus of 1.86. The depositional history with its fast sedimentation along with absent/limited post-mortem transportation led to extraordinarily good preservation of the examined specimen.

Słowa kluczowe

EN

belemnites; Early Cretaceous; Northern Calcareous Alps; phragmocone; proostracum; rostrum; Valanginian

• Wydawca: Faculty of Geology of the University of Warsaw ; Komitet Nauk Geologicznych PAN
• Czasopismo: Acta Geologica Polonica
• Rocznik: 2005
• Tom: Vol. 55, no. 2
• Strony: 147--162
• Opis fizyczny: Bibliogr. 105 poz., rys.

Twórcy

autor

Lukeneder A.

• Natural History Museum Vienna, Department of Geology and Palaeontology, Burgring 7, A-1010, Vienna, Austria

Bibliografia

• ALLISON, P.A. 1988. Phosphatised soft-bodied squids from the Jurassic Oxford Clay. Lethaia, 21, 403-410.
• BARSKOV, I.S. 1972. Microstructure of the skeletal layers of belemnites compared with external shell layers of other mollusks. Paleontological Journal, 1972 (4), 492-500.
• BATHER, F.A. 1888. The growth of Cephalopod shells. Geological Magazine, 4 (3), 44-449.
• BAYLE, E. 1878. Explication de la carte géologique de France 4, 1. Fossiles principaux des terraines. Atlas. Nationale, Paris, 176. pls.
• BLAINVILLE, H.M. D. de 1827. Mémoire Sur Les Bélemnites, Considérées Zoologiquement Et Géologiquement, 136 pp. F.G. Levrault; Paris - Strasbourg.
• BRETT, C.E. & BAIRD, G.C. 1986. Comparative taphonomy: a key to palaeoenvironmental Interpretation based on fossil preservation. Palaios, 1, 207-227.
• BROWN, C. 1900. Über den Genus Hybodus und seine systematische Stellung. Palaeontographica, 46, 149-174.
• CHALLINOR, A.B. 1991. Revision of the belemnites of Misool and a review of the belemnites of Indonesia. Palaeontographica, Abteilung A, 218, 87-164.
• CLÉMENT, A. 1999. Étude paléontologique et biostratigraphique des Duvaliidae (Belemnitida) du Valanginien de la région des Baronnies (bassin vocontien, Sud-Est de la France). Unpublished master thesis, University Claude Bernard, Lyon, 25 pp.
• COMBÉMOREL, R. 1973. Les Duvaliidae Pavlow (Belemnitida) du Crétacé inferieur Français. Documents des laboratoires de géologie de la faculté des sciences de Lyon, 57, 131-185.
• — 1979. Les belemnites. In: R. BUSNARDO, J.-P. THIEULOY & M. MOULLADE (Eds), Hypostratotype mesogéen de l’étage Valanginian (sud-est de la France), Les Stratotypes Français, Comite Francais de Starigraphie, Paris, 6, 69-76.
• — 1988. Lés belemnites de Madagascar. Documents des laboratoires de géologie Lyon, 104, 239 pp.
• COMBÉMOREL, R. & MARIOTTI, M. 1986. Les bélemnites de la carrière de Serra San Quirico (Province d’Ancona, Apennin central, Italie) et la paléobiogéographie des bélemnites de la Téthys méditerranéenne au Tithonique inférieur. Géobios, 19 (3), 299-321.
• COMBÉMOREL, R. & STOYANOVA-VIRGILOVA, M. 1991. Biostratigraphie des belemnites du Crétacé inférieur en France et en Bulgarie. Analyse comparative. Geologica Balcanica, 21, 31-39.
• CUVIER, G. 1795. Mémoire sur la structure interne et externe, et sur les affinities des animaux au quels on a donné le nom de vars. La Décade Philosophique, Litteraire et Politique, 5, 385-396.
• DAUPHIN, Y. 1985. Implications of a microstructural comparison in some fossil and Recent coleoid cephalopod shells. Paläontographica, Abteilung A, 191, 69-83.
• DELATTRE, P.M. 1952. Caractéres et position systematique de Berriasibelus extinctorius (RASPAIL) nov. gen. (Belemnites). Bulletin du Muséum National d’Histoire naturelle, 24, 321-327.
• DOGUZHAEVA, L.A. 2002. Evolutionary trends of Carboniferous coleoids: the ultrastructural view. In: K. WARNKE (Ed.), International Symposium ‘Coleoid cephalopods through time’, Berliner Palöobiologische Abhandlungen, Freie Universität Berlin, Berlin, 1, 29-33.
• DOGUZHAEVA, L.A., MUTVEI, H. & DONOVAN, D.T. 2002a. Structure of the proostracum and muscular mantle in belemnites. 321-339. In: H. SUMMESBERGER, K. HISTON & A. DAURER (Eds), V Inernational Symposium ‘Cephalopods - Present and Past’, 569 pp. Abhandlungen der Geologischen Bundes-Anstalt, 57, Vienna.
• DOGUZHAEVA, L.A., MUTVEI, H. & WEITSCHAT, W. 2002b. Early shell morphology and ultrastructure of Lower Jurassic belemnites from the Ahrensburg erratics (Schleswig Holstein, NW Germany). In: K. WARNKE (Ed.), International Symposium ‘Coleoid cephalopods through time’, Berliner Paläobiologische Abhandlungen, Berlin, 1, 39-42.
• DOGUZHAEVA, L.A., MAPES, R.H. & MUTVEI, H. 2003a. The shell and ink sac morphology and ultrastructure of the Late Pennsylvanian cephalopod Donovaniconus and its phylogenetic significance. In: K. WARNKE, H. KEUPP & S. von BOLETZKY (Eds), International Symposium ‘Coleoid cephalopods through time’, Berliner Paläobiologische Abhandlungen, Berlin, 3, 61-78.
• DOGUZHAEVA, L.A., MUTVEI, H. & WEITSCHAT, W. 2003b. The proostracum and primordial rostrum at the early ontogeny of Lower Jurassic belemnites from north-western Germany. In: K. WARNKE, H. KEUPP & S. von BOLETZKY (Eds), International Symposium ‘Coleoid cephalopods through time’, Berliner Paläobiologische Abhandlungen, Berlin, 3, 79-89.
• DONOVAN, D.T. 1983. Mastigophora OWEN 1856: a little-known genus of Jurassic coleoids. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 484-495.
• DOYLE, P. 1991. Mollusca: Cephalopoda (Coleoidea). In: M.J. BENTOS (Eds), The Fossil Record 2, 229-236. Chapman & Hall, London, Glasgow, New York etc.
• — 1987. Early Cretaceous belemnites from southern Mozambique. Palaeontology, 30, 311-317.
• — 1992. A review of the biogeography of Cretaceous belemnites. Palaeogeography, Palaeoclimatology, Palaeoecology, 92, 207-216.
• DOYLE, P., DONOVAN, D.T. & NIXON, M. 1994. Phylogeny and systematics of the Coleoidea. Palaeontological Bulletin, University of Kansas, new series, 5, 1-15, Lawrence/Kansas.
• DOYLE, P. & MACDONALD, D.I.M. 1993. Belemnite battlefields. Lethaia, 26, 65-80.
• DOYLE, P. & MARIOTTI, N. 1991. Jurassic and Lower Cretaceous belemnites from Northwestern Anatolia (Turkey). In: A. FARINACCI, D.V. AGER & U. NICOSIA (Eds), Geology and paleontology of Western Pontides, Turkey, Geologica Romana, Rome, new Series, 27, 347-379.
• DOYLE, P. & PAGE, K.N. 1994. Belemniten. In: D.M. Martill & J.D. Hudson (Eds), Fossilien aus Ornatenton und Oxford Clay. Ein Bestimmungsatlas, 150-154. Goldschneck-Verlag, Weinstadt-Korb.
• DUVAL-JUVE, J. 1841. Bélemnites des terraines Crétacés inférieurs des environs de Castellane (Basse-Alpes), consid-érées géologiquement et zoologiquement, avec la description de ces terrains. Fortin, Masson et Cie, 80 pp., Paris.
• ENGESER, T. & CLARKE, M.R. 1988. Cephalopod hooks, both Recent and fossil. In: M.R. CLARKE & E.R. TRUEMAN (Eds), The Mollusca, Paleontology and neontology of cephalopods, Academic Press, San Diego, 12, 133-151.
• ENGESER, T. & REITNER, J. 1983. Chitinobelus acifer FISCHER 1981, ein Belemnoteuthide (Coleoidea) mit Epirostrum. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 496-501.
• — & — 1986. Coleoidereste aus der Oberkreide des Libanon im Staatlichen Museum für Naturkunde in Stuttgart. Stuttgarter Beiträge für Natukunde B, 124, 1-15.
• GRAY, J.E. 1849. Catalogue of the Mollusca in the collection of the British Museum. Part 1. Cephalopoda antepedia. VII + 164 pp. Spottiswoodes & Shaw; London.
• GREGORIO, A. de, 1886. Fossiles tithoniques des Stramberg Schichten du “Biancone”, “Roverè di Velo” des Alpes de Verone. Note paléontologique. Annales de Géologie et de Paléontologie, 3, 1-8.
• HAAS, W. 1997. Der Ablauf der Entwicklungsgeschichte der Decabrachia (Cephalopoda, Coleoidea). Palaeontographica Abteilung A, 245, 63-81.
• HEWITT, R.A., LAZELL, B.H. & MOORHOUSE, S.J. 1983. An introduction to the inorganic components of cephalopod shells. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 331-361.
• HEWITT, R.A. & PINCKNEY, G. 1982. On the occurrence and microstructure of the phragmocone and pro-ostracum of the belemnite genus Acroteuthis Stolley. Palaeontographica A, 179, 142-147.
• HEWITT, R.A. & WESTERMANN, G.E.G. 1996. Post-mortem behaviour of early Paleozoic nautilids and paleobathymetry. Paläontologische Zeitschrift, 70, 405-424.
• HEWITT, R.A., WESTERMANN, G.E.G. & JUDD, R.L. 1999. Buoyancy calculations and ecology of Callovian (Jurassic) cylindroteuthid belemnites. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 211 (1/2), 89-112.
• HOEDEMAEKER, P.J., REBOULET, ST., AGUIRRE-URRETA, M., ALSEN, P., AOUTEM, M. ATROPS, F., BARRANGUA R., COMPANY, M., GONZALES, C., KLEIN, J., LUKENEDER, A., PLOCH, I., RAISOSSADAT, N., RAWSON, P., ROPOLO, P., VAŠÍČEK, Z., VERMEULEN, J. & WIPPICH, M. 2003. Report on the 1st International Workshop of the IUGS Lower Cretaceous Ammonite Working Group, the ‘Kilian Group’ (Lyon, 11 September 2002). Cretaceous Research, 24, 89-94.
• HÖLDER, H. 1955. Belemniten und Ammoniten als Beutetiere. Aus der Heimat, 63, 88-92.
• — 1973. Ein belemnitisches Proostracum aus dem Unteren Lias. In: H. HÖLDER (Ed.), Miscellanea cephalopodica. Münsterische Forschungsberichte zur Geologie und Paläontologie, Münster, 29, 63-67.
• HUXLEY, T.H. 1864. On the structure of the Belemnitidae; with a description of a more complete specimen of Belemnites than any hitherto known, and an account of a new genus of Belemnitidae, Xiphoteuthis. Memoirs of the Geological Survey of the United Kingdom, Figures and Descriptions Illustrative of British Organic Remains, 2, 11-36.
• HYATT, A. 1884. Genera of Fosil Cephalopods. Boston Society of Natural History Proceedings, 22, 253-338.
• JANSSEN, N.M.M. 1998. Valanginien-belemniet. Hun verspreiding in ruimte en tijd. Grondboor and Hamer 52: 44-47.
• — 2003. Mediterranean Neocomian belemnites, part 2: the Berriasian-Valanginian boundary in southeast Spain (Río Argos, Cnada Lengua and Tornajo). Scripta Geologica, 126, 121-183.
• JANSSEN, N.M.M. & CLÉMENT A. 2002. Extinction and renewal patterns among Tethyan belemnites in the Verrucosum Subzone (Valanginian) of southeast France. Cretaceous Research, 23, 509-522.
• JANSSEN, N.M.M. & FÖZY, I. 2003. Neocomian belemnites from the Bersek Hill (Gerecse Mountains, Hungary). Földtani Közlöny, 133/2, 291-294.
• JELETZKY, J.A. 1966. Comparative morphology, phylogeny, and classification of fossil Coleoidea. In: R.C. MOORE (Ed.), The University of Kansas Paleontological Contributions, Mollusca, The University of Kansas Publications, Kansas, 7, 162 pp.
• — 1980. Dicoelitid belemnites from the Toarcien - Middle Bajocian of western and Arctic Canada. Geological Survey of Canada, Bulletin, 338, 71 pp.
• KELLER, TH. 1976. Fraßreste im süddeutschen Posidonienschiefer. Jahreshefte der Gesellschaft für Naturkunde in Württemberg, 132, 117-134.
• KOBANOV, G.K. 1967. Skelet belemnitid: Morfologiya i biologicheskie analiz. Trudy Paleontologicheskogo Instituta, 114, 1-100.
• KOSTAK, K., MAREK, J., NEUMANN, P. & PAVELA, M. 2002. An early Carboniferous ?Coleoid (Cephaloppoda, Dibranchiata) fossil from Kulm of Northern Moravia (Czech Republic). In: K. WARNKE (Ed.), International Symposium ‘Coleoid cephalopods through time’, Berliner Paläobiologische Abhandlungen, Berlin, 1, 58-60.
• KRYMGOL’TS, G.YA.1958. Subclass Endocochlia. N.P. LUPPOV & V.V. DRUSCHITS (Eds), Osnovy Paleontologii, MolluskiGeovonogie, Moscow, 2, 145-178. [In Russian]
• LUKENEDER, A. 1999. Acrothoracica-Bohrspuren an einem Belemnitenrostrum (Unterkreide, Obervalanginium; Ober-österreich). Annalen des Naturhistorischen Museums Wien, 101 A, 137-143.
• — 2002. Post-mortem dispersal patterns in lower Cretaceous belemnites (Valanginian, Northern Calcareous Alps, Austria). In: K. WARNKE (Ed.), International Symposium ‘Coleoid cephalopods through time’. Berliner Paläobiologische Abhandlungen Berlin, 1, 68-71.
• — 2004. Late Valanginian ammonoids: Mediterranean and Boreal elements – Implications on sea-level controlled migration (Ebenforst Syncline; Northern Calcareous Alps; Upper Austria. Austrian Journal of Earth Sciences, 95/96, 46-59.
• MANTELL, G.A. 1848. Observations on some belemnites and other fossil remains of cephalopoda, discovered by Mr. Reginald Neville MANTELL, C.E. in the Oxford Clay near Trowbridge, in Wiltshire. Philosophical Transactions of the Royal Society of London, 138, 171-181.
• MONKS, N., HARDWICK, J.D. & GALE, A.S. 1996. The function of the belemnite guard. Paläontologische Zeitschrift, 70, 425-431.
• MUTTERLOSE, J. 1979. Representatives of the subfamily Duvaliinae Pavlow (Belemnitida) from the Hauterivian (Lower Cretaceous) of NW Europe. Aspekte der Kreide Europas. IUGS Series A, 6, 121-127.
• — 1988. Migration and evolution patterns in Upper Jurassic and Lower Cretaceous belemnites. In: J. WIEDMANN & J. KULLMANN (Eds), Cephalopods – Present and Past, pp. 525-537. Schweizerbart’sche Verlagsbuchhandlung; Stuttgart.
• MUTTERLOSE, J., SCHMID, F. & SPAETH, C. 1983. Zur Paläobiogeographie von Belemniten der Unter-Kreide in NWEuropa. Zitteliana, 10, 293-307.
• NAEF, A. 1916. Systematische Übersicht der mediterranen Cephalopoden. Stazione Zoologica di Napoli, Pubblicazioni, 1, 11-19.
• — 1921. Das System der dibranchiaten Cephalopoden und die mediterranen Arten derselben. Zoologische Station zu Neapel, Mitteilungen, 22, 527-542.
• — 1922. Die fossilen Tintenfische. Eine paläozoologische Monographie, 322 pp. Fischer; Jena.
• — 1923. Die Cephalopoden. In: Fauna und Flora des Golfes von Neapel, Monograph Friedländer, Berlin, 35, 149-864.
• NEUMAYR, M. 1873. Die Fauna der Schichten mit Aspidoceras acanthicum. Abhandlungen der kaiserlich königlichen Geologischen Reichsanstalt, 5, 141-257.
• OPPEL, A. 1865. Die Tithonische Etage. Zeitschrift der Deutschen Geologischen Gesellschaft, 17, 535-558.
• OWEN, R.A. 1844. A description of certain belemnites, preserved with a great proportion of their soft-parts, in the Oxford Clay, at Christian Malford - Wiltshire. Philosophical Transactions of the Rroyal Society of London, 1, 65-85.
• ORBIGNY, A.C.V.D. d’ 1855. Paléontologie des coquilles et des mollusques. In: A. DELAHAYS (Ed.), t. 1, 392 pp., Paris.
• PAVLOV, A.P. 1914. Jurassic and Lower Cretaceous cephalopods of northern Siberia. Imperatorskoi Akademii Nauk, St. Petersburg, Zapiski Seriia 8, 21 (4), 68 pp. [In Russian]
• PEARCE, C. 1842. “Concerning Belemnoteuthis”. Geological Society of London, Proceedings, 3, 593 pp.
• POLLARD, J.E. 1968. The gastric contents of an ichthyosaur from the Lower Lias of Lyme Regis, Dorset. Palaeontology, 11, 376-388.
• — 1990. Evidence for diet. In: D.E.G. BRIGGS & P.R. CROWTHER (Eds), Palaeobiology, A Synthesis, 362-367. Blackwell Scientific; Oxford.
• PODLAHA, O.G., MUTTERLOSE, J. & VEIZER, J. 1998. Preservation of δ18O and δ13C in belemnite rostra from the Jurassic/Early Cretaceous successions. American Journal of Science, 298, 324-347.
• RASPAIL, F.V. 1829. Histoire naturelle des Bélemnites, accompagnée de la description et de la classification des espéces que M. Éméric, de Castellane, a recueillies dans les Basses-Alpes de Provence. Annales des Sciences d’Observations, 1, 271-331.
• REITNER, J. & ULRICHS, M. 1983. Echte Weichteilbelemniten aus dem Untertoarcium (Posidonienschiefer) Südwestdeutschlands. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 450-465.
• RIEGRAF, W. 1973. Bißspuren auf jurassischen Belemnitenrostren. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 1973, 494-500.
• — 1977. Fraßreste im süddeutschen Posidonienschiefer. In: TH. KELLER (Ed.), Jahreshefte der Gesellschaft für Naturkunde in Württemberg, Württemberg, 132, 123-126.
• — 1981. Revision der Belemniten des Schwäbischen Jura. Teil 8. Palaeontographica A, 173, 64-139.
• — 1983. Über “Sueviteuthis“ zellensis Reitner and Engeser, 1982 (Coleoidea) aus dem Untertoarcium von Südwestdeutschland. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 502-507.
• — 1987. On Lower and Upper Jurassic dibranchiate cephalopods from Germany and England. Paläontologische Zeitschrift, 64, 261-272.
• — 1995. Cephalopoda dibranchiata fossiles (Coleoidea) [under collaboration of A. Weiss and P. Doyle] In: F. WESTPHAL (Ed.), Fossilium Catalogus. I: Animalia, Kugler, Amsterdam, New York, 133, 411 pp.
• — 1996. Belemniten-Fanghäkchen (Cephalopoda, Coleoidea) aus der Psilonotenbank (Unter Jura, tiefstes Hettangium) von Südwestdeutschland. Stuttgarter Beiträge zur Naturkunde B, 1-38.
• — 1999. Taxonomic status of the belemnite genus Belemnopsis Bayle 1878. Paläontologische Zeitschrift, 73, 59-76.
• RIEGRAF, W. & HAUFF, R.B. 1983. Belemnitenfunde mit Weichkörper, Fangarmen und Gladius aus dem Untertoarcium (Posidonienschiefer) und Unteraalenium (Opalinuston) Südwestdeuschlands. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 466-483.
• RIEGRAF, W., JANSSEN, N.M.M. & SCHMITT-RIEGRAF, C. 1998. Cephalopoda dibranchiata fossiles (Coleoidea) II. In: F. WESTPHAL (Ed.), Fossilium Catalogus Animalia, Backhuys, Leiden, 135, 512 pp.
• RIEGRAF, W. & REITNER, J. 1979. Die “Weichteilbelemniten” des Posidonienschiefers (Untertoarcium) von Holzmaden (Baden-Württemberg) sind Fälschungen. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 1979, 291-304.
• ROGER, J. 1952. Sous-classe des Dibranchiata Owen, 1836. (Coleoidea WAAGEN, Endocochlia SCHWARZ). 689-755. In: J. PIVETEAU (Ed.), “Traité de paléontologie”, 2, 790 pp. Masson et Cie. Edit; Paris.
• SCHLEGELMILCH, R. 1998. Die Belemniten des Süddeutschen Jura, 155 pp. Gustav Fischer Verlag; Stuttgart – Jena – Lübeck – Ulm.
• SEIBERTZ, E. & SPAETH, C. 2002. Cretaceous belemnites of Mexico. III. The Albian Neo- and Mesohibolites of the “Mexican Solnhofen” Tepexi de Rodriguez (State of Puebla) and their biostrationomy (Lower Cretaceous). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 225, 55-74.
• SEILACHER, A. 1983. Papers on cephalopod paleobiology and phylogeny (editorial introduction). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 165, 327-329.
• SPAETH, C. 1975. Zur Frage der Schwimmverhältnisse bei Belemniten in Abhängigkeit vom Primärgefüge der Hartteile. Paläontologische Zeitschrift, 49, 321-331.
• STEVENS, G.R. 1965. The Jurassic and Cretaceous belemnites of New Zealand and a review of the Jurassic and Cretaceous Belemnites of the indo-Pacific region. New Zealand. New Zealand Geological Survey Palaeontological, Bulletin, 36, 1-283.
• STOLLEY, E. 1919. Die Systematik der Belemniten. Jahresbericht des Niedersächsischen Geologischen Vereins, 11, 1-59.
• STOYANOVA-VERGILOVA, M. 1965. Representants de la SousFamille Duvaliinae Pavlow (Belemnitida) du Cretace Inferieur en Bulgarie. Travaux sur la Geologie de Bulgarie, Serie Paleontologie, 7, 179-221.
• — 1970. Les fossils de Bulgarie. Iva Crétacé infeérieur. Belemnitida, 72 pp. Académie Bulgare des Sciences.
• TOUCAS, A. 1890. Étude de la faune des couches tithonique de l’Ardèche. Bulletin de la Société Géologique de France (3), 18, 560-630.
• VOLTZ, M. 1830. Observations sur les belemnites. Memoires de la Societe du Museum d’Histoire Naturelle de Strasbourg, 1, 1-70.
• WEISS, A.F. 1991. Krevizii belemnitov roda Conobelus STOLLEY, 1919. Paleontologičeskij Zhurnal, 1991 (2), 18-31. [In Russian]
• — 1992. Revision of the belemnite genus Conobelus STOLLEY, 1919. Paleontologičeskij Zhurnal, 25, 21-40.
• WESTERMANN, G.E.G. 1985. Post-mortem descent with septal implosion in Silurian nautiloids. Paläontologische Zeitschrift, 59, 79-97.
• ZITTEL, K.A. 1870. Die Fauna der aelteren Cephalopodefuehrenden Tithonbildungen. Palaeontographica, Supplement, 2, 1-192.