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Ammonites of the Pachydiscus neubergicus group from the mid-Maastrichtian chalk of Denmark: alternative taxonomic interpretations

Machalski Marcin, Owocki Krzysztof, Rasmussen Jan Audun, Madsen Henrik

Acta Geologica Polonica

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2026

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Vol. 76, no. 1

art. no. e70

EN

Despite their rarity, ammonites of the Pachydiscus neubergicus group rank amongst the stratigraphically most important macrofossils in the Maastrichtian of Europe. Here, their taxonomy is studied on the basis of material from mid-Maastrichtian chalk exposed at Hillerslev in Denmark. The studied sample consists of ten specimens in mouldic preservation, collected from a c. 10 m thick chalk interval. Their taxonomic identification is based on comparison with ‘type populations’ of known members of the P. neubergicus group, primarily with the roughly coeval material of P. neubergicus (v. Hauer, 1858) from Neuberg in Austria. To date, only P. neubergicus has been identified from this group in the Danish chalk. Based largely on differences in ribbing density, the Hillerslev material has conventionally been subdivided into several taxa, namely P. neubergicus, P. sp. cf. neubergicus, P. jacquoti (Seunes, 1890), P. sp. cf. jacquoti, P. sp. aff. armenicus Atabekian and Akopian, 1969, and Pachydiscus sp. Results of the Principal Component Analysis (PCA) and one-sample t test for the studied material are compatible with such identifications of the Hillerslev specimens. However, an alternative taxonomic interpretation is possible that the identified morphospecies, with the potential exception of P. sp. aff. armenicus, are members of a single, highly-variable biospecies, dominated by a sparsicostate morphotype closer to P. jacquoti than P. neubergicus. Additional material from the mid-Maastrichtian levels of Danish chalk is needed to test the latter hypothesis.

2

Suevisphinctes – a new perisphinctid ammonite genus from the Lower Tithonian (Hybonotum Zone) of Southern Germany

Scherzinger Armin, Schweigert Günter, Parent Horacio, Főzy István

Volumina Jurassica

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2024

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Vol. 22

13--24

EN

A new Upper Jurassic perisphinctid ammonite genus, Suevisphinctes (type species: Suevisphinctes josefkelleri nov. gen., nov. sp.), is established. At present, it is recorded from the laisackerensis biohorizon (Early Tithonian, Hybonotum Zone, Rueppellianus Subzone) of SW Germany. A historical ammonite specimen of supposed Tithonian age from the Polish Klippen of Iskrzyczyn may be assigned to Suevisphinctes as well and thus indicates a possible Tethyan origin of this enigmatic genus.

3

On some Turonian and Coniacian ammonites from the Czech Republic

Kennedy William James, Summesberger Herbert

Acta Geologica Polonica

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2024

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Vol. 74, no. 2

art. no. e10

EN

Ammonites, chiefly from the Upper Turonian Prionocyclus germari Zone, the Lower Coniacian part of the Forresteria (Harleites) petrocoriensis Zone, and the Lower Coniacan Peroniceras (Peroniceras) tridorsatum Zone from the Czech Republic are figured and described, including classic material originally described by Fritsch (1872–1898) and Jahn (1892, 1896). In all, 25 species are recognised, including two, Pseudojacobites sp. and Muniericeras sp., that represent genera previously not known from the area.

4

The ammonite subgenus Pervinquieria (Deiradoceras) van Hoepen, 1931 from the Upper Albian of KwaZulu-Natal, South Africa. Part I

Kennedy William James, Klinger Herbert Christian

Acta Geologica Polonica

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2023

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Vol. 73, no. 4

549--569

EN

The cosmopolitan Late Albian ammonite subgenus Pervinquieria (Deiradoceras) van Hoepen, 1931, and its synonyms Cechenoceras van Hoepen, 1941 and Mimeloceras van Hoepen, 1944, originally based on material from northern KwaZulu-Natal, are reviewed. The type material of the type species, Subschloenbachia prerostrata Spath, 1921, is revised and reillustrated, as are its numerous synonyms.

5

The ammonite subgenus Pervinquieria (Deiradoceras) van Hoepen, 1931 from the Upper Albian of KwaZulu-Natal, South Africa. Part II

Kennedy William James, Klinger Herbert Christian

Acta Geologica Polonica

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2023

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Vol. 73, no. 4

571–-586

EN

In this contribution we complete the revision of species we refer to Pervinquieria (Deiradoceras) van Hoepen, 1931, focusing on those assigned by him to his genera Cechenoceras and Mimeloceras.

6

The ammonite genera Dipoloceras, Diplasioceras, Euspectroceras, and Rhytidoceras from the Upper Albian of KwaZulu-Natal, South Africa

Kennedy William James, Klinger Herbert Christian

Acta Geologica Polonica

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2023

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Vol. 73, no. 4

505–-548

EN

The lower Upper Albian ammonite genus Dipoloceras Hyatt, 1900 (subfamily Mojsisovicsiinae Hyatt, 1903) is represented in KwaZulu-Natal by the type species, D. cristatum (Brongniart, 1822), marker species for the base of the Upper Albian Substage. Rhytidoceras van Hoepen, 1931 (of which Drepanoceras van Hoepen, 1931, non Stein 1878 and Ricnoceras van Hoepen, 1941, are synonyms), previously regarded as subgenera of Dipoloceras, are afforded generic status as are its supposed synonyms Diplasioceras van Hoepen, 1946a, and Euspectroceras van Hoepen, 1946a. The type species of these genera are revised, and assigned to the subfamily Pervinquierinae Spath, 1926.

7

Stratigraphically significant, sporadic Early Cretaceous ammonites in Butkov Quarry (Central Western Carpathians, Slovakia)

Vašíček Zdeněk

Acta Geologica Polonica

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2023

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Vol. 73, no. 4

587–-612

EN

Butkov Quarry provides the best exposed stratigraphic sequence of marly limestones with Early Cretaceous ammonites in the Manín Nappe of the Central Western Carpathians. The presented paper deals with the sporadically occurring zonal ammonites, or ammonites of guiding character, from the Lower Valanginian to Upper Hauterivian. Sixteen species are taxonomically elaborated here in detail. More attention is given to the basic taxonomy of the Subfamily Crioceratitinae Gill, 1871. The species described here, like most of the previously published species from Butkov Quarry, are representatives of the Mediterranean bioprovince and are close to the ammonite association from the Vocontian Basin.

8

On some Upper Cretaceous ammonites from western Uzbekistan

Ward David J., King Chris, Morris Noel J., Kennedy William James

Acta Geologica Polonica

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2023

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Vol. 73, no. 4

613–-634

EN

Upper Cretaceous ammonites are described from six horizons in the mammal-bearing terrigenous-clastic sequences of western Uzbekistan, as follows: Upper Cenomanian, with Placenticeras sp. juv. cf. cumminsi Cragin, 1893 and Metoicoceras geslinianum (d’Orbigny, 1850); Lower Turonian with Tragodesmoceras cf. mauryae Kennedy and Wright, 1981, Placenticeras kharesmense (Lahusen, 1884), Watinoceras coloradoense (Henderson, 1908), Watinoceras amudariense (Arkhangelsky, 1916b), Metasigaloceras rusticum (J. Sowerby, 1823), Morrowites wingi (Morrow, 1935), Sciponoceras cf. bohemicum bohemicum (Fritsch, 1872), and Yezoites amudariensis (Arkhangelsky, 1916b); Middle Turonian with Collignoniceras woollgari woollgari (Mantell, 1822); Upper Turonian with Lewesiceras mantelli (Wright and Wright, 1951) and Placenticeras crassum Ilyin, 2020; a single Placenticeras semiornatum (d’Orbigny, 1850) from around the Coniacian/Santonian boundary, and Santonian Placenticeras polyopsis (Dujardin, 1837).

9

Phylogeny of the ammonite family Aulacostephanidae Spath, 1924 during the Late Oxfordian and the Early Kimmeridgian in Europe: Main lineages, patterns of evolution and sedimentological to palaeogeographical controls on evolutionary development

Wierzbowski Andrzej

Volumina Jurassica

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2022

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Vol. 20

59--128

EN

The evolutionary history of the Aulacostephanidae presented here includes the interval of about 5 Ma of the Late Oxfordian and Early Kimmeridgian ranging from the origin of the family to is maximal development. The development and biogeographical distribution of aulacostephanids in Europe were related mostly to sea-level changes and tectonics. The appearance of the ancestor genus Decipia and the following genus Ringsteadia was controlled by marine transgressions during the Late Oxfordian which stimulated also the distribution of these ammonites in Subboreal and Submediterranean Europe. The main faunal turnover at the Oxfordian/Kimmeridgian boundary was related to allopatric speciation correlated to separation of the aulacostephanid assemblages: it resulted in the appearance of the main Ringsteadia-Pictonia-Rasenia lineage in NW Subboreal Europe and the side-lineages Ringsteadia-Vielunia-Eurasenia/Involuticeras and Ringsteadia-Vineta-Balticeras-Rasenioides in NE Subboreal and Central Submediterranean Europe. A strong development of the shallow-water carbonate platforms during the Early Kimmeridgian stimulated the development of still another Pictonia (Pomerania) - Rasenia (Pachypictonia) lineage. The main faunal turnover correlated to a large transgression at the end of the Early Kimmeridgian resulted in the sudden distribution of the new aulacostephanid faunas over large areas of northern and central Europe, and opened a new stage in the evolutionary development of the family.

10

Upper upper Albian (Mortoniceras rostratum Zone) cephalopods from Clansayes (Drôme, south-eastern France)

Jattiot Romain, Lehmann Jens, Latutrie Benjamin, Tajika Amane, Vennin Emmanuelle, Vuarin Pauline, Brayard Arnaud, Fara Emmanuel, Trincal Vincent

Acta Geologica Polonica

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2022

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Vol. 72, no. 2

187--233

EN

We document an upper upper Albian (Mortoniceras rostratum Zone) cephalopod assemblage from Clansayes (Drôme, south-eastern France). Although fossils are rare in local exposures and in the single sampled level, a decade of intensive fossil collecting yielded 290 ammonite and 5 nautilid specimens. In total, we describe 1 species of nautilid and 24 species (within 17 genera) of ammonites, including 13 heteromorphs. Only two of these ammonite taxa were previously recorded from the upper upper Albian at Clansayes, which demonstrates the value of this fauna with regard to taxonomy, palaeobiology and palaeobiogeography. Based on morphological and biometric analyses performed on an extensive material (104 specimens), we discriminate two species for the heteromorphic ammonite genus Mariella Nowak, 1916 within the Mortoniceras rostratum Zone. In addition, we investigate shell chirality patterns in Mariella from the late Albian of southern France. Upon comparison of the Clansayes material with older material from the immediately underlying upper Albian Mortoniceras fallax Zone at the neighbouring Salazac locality, we identify an increase in the proportion of sinistral specimens. This observed increase in the frequency of sinistral Mariella specimens may hypothetically be part of a global evolutionary pattern, considering that nearly all documented younger Cenomanian Mariella (and more generally Cenomanian turrilitids) are sinistral.

11

New records of Tmaegoceras lacordarii (Michelin, 1835) from the Lower Jurassic Arietenkalk Formation of SW Germany and its systematic relationships

Schweigert Günter, Scherzinger Armin, Gräbenstein Stefan

Volumina Jurassica

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2021

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Vol. 19, no. 1

95--108

EN

New material of the enigmatic cosmopolitan ammonite genus Tmaegoceras Hyatt, 1889 is reported from the upper Lower Sinemurian (Bucklandi Zone, ‛cf. pinguisʼ Biohorizon) of SW Germany. It is better preserved than previously described material. All specimens are assigned to Tmaegoceras lacordarii (Michelin, 1835), an almost forgotten subjective senior synonym of Tmaegoceras crassiceps Pompeckj, 1901. The presence of an extreme rursiradiate ribbing style in some specimens allows a better understanding of this genus and provides a hint for its systematic affiliation. This diagnostic ribbing style is shared with some extremely rare Alpine taxa such as Ammonites salinarius Hauer, 1846 and “Arietites” subsalinarius Wähner, 1891. Tmaegoceras is included here in Pseudotropitinae Donovan, 1973. Pseudotropitinae are possibly an early offshoot of Arietitidae indigenous of the Panthalassa Realm that occasionally spread into the Tethys and its adjacent shelves. A phyletic relationship with Late Triassic Tropitidae Mojsisovics, 1875, however, cannot be excluded either.

12

The microcrinoid taxonomy, biostratigraphy and correlation of the upper Fredericksburg and lower Washita groups (Cretaceous, middle Albian to lower Cenomanian) of northern Texas and southern Oklahoma, USA

Gale Andrew Scott, Rashall Jenny Marie, Kennedy William James, Holterhoff Frank Koch

Acta Geologica Polonica

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2021

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Vol. 71, no. 1

1--52

EN

The stratigraphy of the upper Fredericksburg and lower Washita groups of northern Texas and southern Oklahoma is described, and biostratigraphical correlation within the region, and further afield, using micro-crinoids, ammonites, planktonic foraminiferans and inoceramid bivalves is summarised. The taxonomy of the roveacrind microcrinoids is revised by the senior author, and a new genus, Peckicrinus, is described, with the type species Poecilocrinus porcatus (Peck, 1943). New species include Roveacrinus proteus sp. nov., R. morganae sp. nov., Plotocrinus reidi sp. nov., Pl. molineuxae sp. nov., Pl. rashallae sp. nov. and Styracocrinus thomasae sp. nov. New formae of the genus Poecilocrinus Peck, 1943 are Po. dispandus forma floriformis nov. and Po. dispandus forma discus nov. New formae of the genus Euglyphocrinus Gale, 2019 are E. pyramidalis(Peck, 1943) forma pyramidalis nov., E. pyramidalis forma radix nov. and E. pyramidalis forma pentaspinusnov. The genera Plotocrinus Peck, 1943, Poecilocrinus and Roveacrinus Douglas, 1908 form a branching phylogenetic lineage extending from the middle Albian into the lower Cenomanian, showing rapid speciation, upon which a new roveacrinid zonation for the middle and upper Albian (zones AlR1-12) is largely based. Outside Texas and Oklahoma, zone AlR1 is recorded from the lower middle Albian of Aube (southeastern France) and zones AlR11-CeR2 from the Agadir Basin in Morocco and central Tunisia. It is likely that the zonation will be widely applicable to the middle and upper Albian and lower Cenomanian successions of many other regions.

13

Stratigraphy of the Albian-Cenomanian boundary interval in the Agadir Basin, Morocco: ammonites, microcrinoids, planktonic foraminifera

Gale Andrew Scott, Kennedy William James, Petrizzo Maria Rose

Acta Geologica Polonica

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2021

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Vol. 71, no. 4

453--480

EN

The uppermost Albian and lowermost Cenomanian succession at Abouda Plage, north of Agadir, in the Agadir Basin, western Morocco, is described in detail, and ammonites, microcrinoids and planktonic foraminifera are recorded and illustrated. The lower part of the Aït Lamine Formation yields ammonites indicative of the Pervinquieria (Subschloenbachia) rostrata and P. (S.) perinflata ammonite zones, and the Thalmanninella appenninica planktonic foraminiferan Zone. The base of the Cenomanian is identified at 42.2 m above the base of the Aït Lamine Formation, based on the lowest occurrence of the planktonic foraminiferan Thalmanninella globotruncanoides Sigal, 1948. Lower Cenomanian ammonites of the Graysonites adkinsi Zone enter 3 m higher in the succession. Microcrinoid zones AlR11 and AlR12 are identified in the Upper Albian, and the base of the CeR1 Zone coincides with the lowest occurrence of Cenomanian ammonites. The ammonite and microcrinoid occurrences and detailed distributions are very similar to those found in north central Texas, which, in the Cenomanian, was 5,300 km to the west. The new records suggest that the G. adkinsi Zone is equivalent to the uppermost (Lower Cenomanian) part of the Pleurohoplites briacensis Zone of the Global Stratotype Section for the base of the Cenomanian stage. An hiatus, of global extent, immediately underlies the base of the G. adkinsi Zone and is represented in the Agadir Basin by an erosion surface containing bored and encrusted hiatus concretions.

14

On some Turonian and Coniacian ammonites from central Colombia

Kennedy William James

Acta Geologica Polonica

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2021

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Vol. 71, no. 3

259--285

EN

The rich collections of Turonian and Coniacian ammonites from Colombia, housed in the University of California Museum of Paleontology, Berkeley, are described, with 20 species recognized, referred to the genera Reymenticoceras Kennedy, 2018, Vascoceras Choffat, 1898, Fagesia Pervinquière, 1907, Wrightoceras Reyment, 1954a, Subprionocyclus Shimizu, 1932, Subprionotropis Basse, 1951, Prionocycloceras Spath, 1926, Barroisiceras de Grossouvre, 1894, Forresteria Reeside, 1932, Niceforoceras Basse, 1948, Peroniceras de Grossouvre, 1894, Gauthiericeras de Grossouvre, 1894, Protexanites Matsumoto, 1955, and Metaptychoceras Spath, 1926. One species, Reymenticoceras ornatum, is new. Most of the species described can be assigned to the substages and ammonite zones recognized in Western Europe.

15

On imperfectly known Hauterivian representatives of the families Holcodiscidae Spath, 1923 and Barremitidae Breskovski, 1977 in Butkov Quarry (Central Western Carpathians, Slovakia)

Vašíček Zdeněk, Klein Jaap

Acta Geologica Polonica

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2021

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Vol. 71, no. 4

433--451

EN

In Butkov Quarry, ammonites of the families Holcodiscidae Spath, 1923 and Barremitidae Breskovski, 1977 occur in the pelagic Lower Cretaceous pelagic deposits of the Manín Unit. This contribution discusses the taxonomy of both families and presents their distribution in the layered sequences of the quarry. The genus Spitidiscus Kilian, 1910 classified as a member of the Superfamily Perisphinctoidea Steinmann in Steinmann and Döderlein, 1890 is an important representative of the Holcodiscidae from a stratigraphic point of view. In areas where the zonal index Acanthodiscus radiatus (Bruguière, 1789) does not occur, as in Butkov Quarry, the first representatives of Spitidiscus indicate the base of the Hauterivian. The genus Plesiospitidiscus Breistroffer, 1947 was long regarded as a member of the Superfamily Desmoceratoidea Zittel, 1895. This superfamily was based on its type species, Eodesmoceras celestini (Pictet and Campiche, 1860), which is not Valanginian in age, as now clearly proven. As a consequence, this superfamily is considered invalid. Vermeulen and Lahondère (2011) proposed an alternative by selecting a suitable initial genus, namely Plesiospitidiscus, for the Family Barremitidae, Superfamily Barremitoidea Breskovski, 1977 (nom. transl. Vermeulen and Lahondère, 2011).

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The Bajocian to Kimmeridgian (Middle to Upper Jurassic) ammonite succession at Sentralbanken High (core 7533/3-U-1), Barents Sea, and its stratigraphical and palaeobiogeographical significance

Andrzej Wierzbowski, Morten Smelror

Volumina Jurassica

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2020

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Vol. 18, no. 1

1--22

EN

Ammonites recovered from Upper Bajocian to Upper Kimmeridgian strata in a core drilled at Sentralbanken High in the northern Barents Sea are described and chronostratigraphically interpreted. The lower part of the core comprises Upper Bajocian to Upper Callovian deposits of the Fuglen Formation with ammonites of the genus Cranocephalites near the base, and Longaeviceras in the upper part. The overlying Oxfordian to Kimmeridgian deposits of the Hekkingen Formation are thin and condensed. The Hekkingen Formation is dominated by Boreal cardioceratid ammonites (Cardioceras, Amoebites, Euprionoceras, Hoplocardioceras), with numerous Subboreal aulacostephanid ammonites (Rasenia, Zenostephanus) at two levels. The occurrences of Boreal and Subboreal ammonites are discussed in relation to the palaeogeography and fluctuations of ammonite faunas within the Boreal Realm during the Kimmeridgian with special attention to levels rich in Subboreal ammonites whose appearance has been controlled by tectonic and climatic factors. The uppermost part of the succession deposited during a time of maximum flooding in the Late Kimmeridgian is dominated by Boreal ammonites.

17

Hypowaagenia nov. gen., a rare genus of giant aspidoceratid ammonite from the Upper Jurassic of Southern Germany

Schweigert Günter, Schlampp Victor

Volumina Jurassica

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2020

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Vol. 18, no. 1

23--36

EN

A new genus, Hypowaagenia (type species: H. endressi nov. sp.), is introduced for previously unknown large-sized macroconchiate aspidoceratid ammonites with an umbilical and a ventrolateral row of spines. In the medium and adult stages irregular ribs interconnect these spines. The oldest unequivocal record of this genus comes from the Early Kimmeridgian Planula Zone, whereas the type material of H. endressi nov. sp. comes from the upper Platynota Zone/? lowermost Hypselocyclum Zone. Corresponding microconchs have not yet been identified. Although all unequivocal records come from Southern Germany, a Tethyan origin is most likely for these aspidoceratids. Another, stratigraphically younger species tentatively included in Hypowaagenia is Aspidoceras acanthomphalum (Zittel, 1870), which is, however, only recorded yet by mesoconchiate specimens. The ammonite fauna of the type horizon of H. endressi nov. sp. is briefly characterized and termed as the geniculatum Biohorizon of the late Platynota Zone.

18

Arietites solarium (Quenstedt, 1883) – a diagnostic ammonite species in the Lower Jurassic (Early Sinemurian, Bucklandi Zone) of SW Germany

Scherzinger Armin, Gräbenstein Stefan, Schweigert Günter

Volumina Jurassica

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2020

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Vol. 18, no. 1

37--46

EN

The coarse-ribbed and big-sized Arietites solarium (Quenstedt, 1883) is one of the largest-grown and most iconic ammonite taxa in the entire Swabian Lower Jurassic; however, despite previous revisions including the designation of a lectotype, there has been some confusion concerning its correct identification, and its type horizon within the Lower Sinemurian Arietenkalk Formation was not exactly known. Arietites solarium characterises the herein introduced solarium Biohorizon of the upper Bucklandi Zone of the Sinemurian. For nomenclatorial stability, we designate a neotype based on the only surviving specimen of Quenstedtʼs original type series. In addition, we provide a preliminary succession of recognized biohorizons in the Lower Sinemurian of Swabia, which will make correlations with other areas more reliable.

19

A microconchiate Hypowaagenia Schweigert and Schlampp, 2020 (Aspidoceratidae, Hybonoticeratinae) from the Upper Jurassic of Southern Germany

Schweigert Günter, Schlampp Victor

Volumina Jurassica

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2020

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Vol. 18, no. 2

115--120

EN

The previously unknown microconch which corresponds to the recently introduced Late Jurassic aspidoceratoid ammonite genus Hypowaagenia Schweigert and Schlampp, 2020, is reported from beds of the topmost Platynota Zone or basal Hypselocyclum Zone of Franconia. This record indicates that these exotic ammonite findings are not of long-drifting necroplanktonic shells, but stem from animals that have spread over this area after immigration from the Tethys

20

Upper Albian, Cenomanian and Upper Turonian ammonite faunas from the Fahdène Formation of Central Tunisia and correlatives in northern Algeria

Kennedy William James

Acta Geologica Polonica

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2020

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Vol. 70, no. 2

147--272

EN

Over 130 species are documented from the Upper Albian, Cenomanian and Upper Turonian Fahdène Formation and correlatives in Central Tunisia and northern Algeria, based on material described by Henri Coquand (1852, 1854, 1862, 1880), Léon Pervinquière (1907, 1910), Georges Dubourdieu (1953), Jacques Sornay (1955), and new collections. The material consists predominantly of limonitic nuclei, together with adults of micromorphs. There is no continuous record, and a series of faunas are recognised that can be correlated with the zonation developed in Western Europe. These are the Upper Albian Ostlingoceras puzosianum fauna, Lower Cenomanian Neostlingoceras carcitanense and Mariella (Mariella) harchaensis faunas, the upper Lower to lower Middle Cenomanian Turrilites scheuchzerianus fauna, Middle Cenomanian Calycoceras (Newboldiceras) asiaticum fauna, Upper Cenomanian Eucalycoceras pentagonum fauna, and the Upper Turonian Subprionocyclus neptuni fauna. Two new micromorph genera are described, Coquandiceras of the Mantelliceratinae and Cryptoturrilites of the Turrilitinae. Most of the taxa present have a cosmopolitan distribution, with a minority of Boreal, North American and endemic taxa.